形態特征
由于(yu)(yu)當地群(qun)眾有(you)燒(shao)墾的(de)(de)(de)習慣,自然(ran)植被多(duo)被燒(shao)毀,分(fen)布范圍(wei)狹窄(zhai),加以(yi)本種(zhong)(zhong)(zhong)開花結(jie)實(shi)的(de)(de)(de)周期(qi)長,天然(ran)更(geng)新能(neng)力弱,1987年(nian)(nian)被列為(wei)世界最(zui)瀕危的(de)(de)(de)12種(zhong)(zhong)(zhong)植物之一(yi)(yi)(yi)。日(ri)前(qian),在(zai)百(bai)山(shan)(shan)祖國家級自然(ran)保護區內,80多(duo)株百(bai)山(shan)(shan)祖冷(leng)杉種(zhong)(zhong)(zhong)子(zi)實(shi)生(sheng)苗正茁(zhuo)壯成(cheng)長。這是繼1992年(nian)(nian)后(hou),時隔20年(nian)(nian)又一(yi)(yi)(yi)次成(cheng)功(gong)繁育(yu)的(de)(de)(de)原生(sheng)樹種(zhong)(zhong)(zhong)子(zi)幼苗。百(bai)山(shan)(shan)祖冷(leng)杉的(de)(de)(de)再次成(cheng)功(gong)育(yu)苗對(dui)于(yu)(yu)保護拯救瀕臨(lin)滅絕的(de)(de)(de)這一(yi)(yi)(yi)物種(zhong)(zhong)(zhong)具有(you)深(shen)遠意義。喬(qiao)木(mu),高約11米,胸徑40厘米;樹皮(pi)(pi)灰(hui)(hui)白色,不規則(ze)(ze)龜裂(lie)(lie),裂(lie)(lie)塊縱向(xiang)大于(yu)(yu)橫(heng)向(xiang),裂(lie)(lie)隙(xi)較(jiao)深(shen);大枝(zhi)平(ping)展,枝(zhi)皮(pi)(pi)不規則(ze)(ze)淺裂(lie)(lie);小(xiao)枝(zhi)對(dui)生(sheng),稀三枝(zhi)輪生(sheng),基部圍(wei)有(you)宿存芽(ya)(ya)鱗,主(zhu)干及直立枝(zhi)上的(de)(de)(de)小(xiao)枝(zhi)交(jiao)叉對(dui)生(sheng),一(yi)(yi)(yi)年(nian)(nian)生(sheng)枝(zhi)淡黃(huang)色或黃(huang)灰(hui)(hui)色,無毛或凹槽中有(you)疏毛;冬(dong)芽(ya)(ya)卵圓形,有(you)樹脂,生(sheng)于(yu)(yu)枝(zhi)頂,三個排成(cheng)一(yi)(yi)(yi)平(ping)面(mian),中間(jian)之芽(ya)(ya)常(chang)較(jiao),兩側之芽(ya)(ya)為(wei)大,稀在(zai)小(xiao)枝(zhi)節(jie)間(jian)上面(mian)(即背地面(mian))生(sheng)有(you)一(yi)(yi)(yi)芽(ya)(ya)、其后(hou)發(fa)育(yu)成(cheng)直立枝(zhi),芽(ya)(ya)鱗淡黃(huang)褐(he)色,三角狀卵形,背面(mian)中上部具鈍縱脊(ji)。
葉條形(xing),在(zai)側枝上排列(lie)(lie)成二列(lie)(lie)狀(zhuang),或(huo)枝條下面(mian)(mian)(mian)之(zhi)(zhi)葉排列(lie)(lie)成兩(liang)(liang)列(lie)(lie),上面(mian)(mian)(mian)之(zhi)(zhi)葉斜展至直伸,其長度由下面(mian)(mian)(mian)兩(liang)(liang)側至上面(mian)(mian)(mian)中央遞(di)減(jian),長1-4.2(通常為1.5-3.5)厘(li)米(mi),寬2.5-3.5毫米(mi),先端(duan)(duan)有(you)(you)凹缺,下面(mian)(mian)(mian)有(you)(you)兩(liang)(liang)條白色氣孔帶;橫切面(mian)(mian)(mian)有(you)(you)2個邊(bian)生樹脂(zhi)道(dao)或(huo)生于兩(liang)(liang)側端(duan)(duan)的葉肉薄壁組織內(nei);上面(mian)(mian)(mian)至下面(mian)(mian)(mian)兩(liang)(liang)端(duan)(duan)及下面(mian)(mian)(mian)中部有(you)(you)一層連續(xu)排列(lie)(lie)的皮下層細胞;幼枝之(zhi)(zhi)葉在(zai)枝上常呈兩(liang)(liang)列(lie)(lie),先端(duan)(duan)呈二尖裂。
雌(ci)球(qiu)花(hua)(hua)圓柱(zhu)形(xing)(xing),長(chang)(chang)3-3.5厘(li)米,徑約8毫米,苞鱗上(shang)部向后反曲。球(qiu)果通常(chang)每一枝節之(zhi)(zhi)間(jian)著生1-3個,圓柱(zhu)形(xing)(xing),長(chang)(chang)7-12厘(li)米,徑3.5-4厘(li)米,成熟(shu)前綠色至淡黃綠色,熟(shu)后淡褐(he)黃色或淡褐(he)色;中部種四邊(bian)(bian)(bian)形(xing)(xing),稀近腎狀四邊(bian)(bian)(bian)形(xing)(xing),長(chang)(chang)1.8-2.4厘(li)米,寬(kuan)2.5-3厘(li)米,先(xian)端近全緣或有極細之(zhi)(zhi)細齒(chi),兩側邊(bian)(bian)(bian)緣有不規(gui)則鋸齒(chi),基部楔(xie)形(xing)(xing)、兩側耳狀;苞鱗稍短于(yu)種鱗或幾(ji)相等(deng)長(chang)(chang),上(shang)部近圓形(xing)(xing),寬(kuan)7-8毫米,邊(bian)(bian)(bian)緣有細齒(chi),先(xian)端露(lu)出、反曲,尖頭短,長(chang)(chang)1毫米以內,中部收縮或窄縮呈條狀,寬(kuan)3.5-4毫米;種子呈倒三角(jiao)狀,具與種子等(deng)長(chang)(chang)而寬(kuan)大(da)的膜質種翅,翅端平截。花(hua)(hua)期熟(shu)。
生長環境
百山(shan)祖冷杉產(chan)地(di)位于東部亞熱(re)帶高(gao)山(shan)地(di)區,氣候特點是溫(wen)(wen)度(du)極(ji)低,濕度(du)大(da),降(jiang)水(shui)多,云霧重。年平均溫(wen)(wen)度(du)8~9℃,極(ji)端最低-15℃;年降(jiang)水(shui)量達2 300毫米(mi),相對濕度(du)92%。成土(tu)母質多為凝灰 巖、流紋巖之(zhi)風化物(wu),土(tu)壤為黃棕(zong)壤,呈(cheng)酸性(xing),pH值4.5,有(you)機質含量3.5%。
自然植被(bei)為(wei)落葉闊葉林(lin),伴(ban)生植物主要(yao)有(you)亮葉水青(qing)岡 ,林(lin)下木(mu)為(wei)百山祖玉山竹(zhu)和華赤竹(zhu)。本(ben)種(zhong)幼樹極耐陰,但生長不良。大樹枝條常向光(guang)面(mian)屈曲。結(jie)實周期4-5年,多數種(zhong)子(zi)發(fa)育不良,5月開(kai)花,11月球果成熟。
分布范圍
為中(zhong)國東南(nan)部新近發現(xian)的稀有珍貴樹(shu)種,特產于浙江南(nan)部百(bai)山祖南(nan)坡(po)海拔1700米以上地帶,林中(zhong)混生的優勢樹(shu)種為亮葉(xie)山毛櫸及多脈(mo)青岡櫟(li)、青岡櫟(li)等(deng)闊葉(xie)樹(shu)種。
繁殖方法
嫁接繁殖:
①在(zai)雌株上嫁接有雄花的枝條(tiao);
②以日本冷杉為砧木(mu)的靠(kao)接插,成活后立即分開(kai),以減少砧木(mu)對接穗的影(ying)響。
生物工程新技(ji)術(shu)的應用
①莖尖培養(yang),國(guo)外已獲成功,可以應(ying)用;
②種胚培養;
③花藥培養;
④根尖或愈傷(shang)組織(zhi)培養。
為了(le)迅速繁殖百山祖冷杉(shan),1977年(nian)以來,連續進行無性繁殖試驗。1975年(nian)采集了(le)一批百山祖冷杉(shan)種(zhong)(zhong)子,但1976年(nian)播種(zhong)(zhong)后不見發芽,說明這批種(zhong)(zhong)子是無效的,僅有種(zhong)(zhong)子的外型,而沒有胚的發育。1977年(nian)3月(yue)29日,第一次進行扦(qian)插試驗,但未獲成功。
1978年(nian)用(yong)磷酸二氫鉀液泡(pao)插穗(sui),進(jin)行扦(qian)(qian)插,因管(guan)理(li)不(bu)善失敗(bai),對照的(de)日(ri)(ri)本冷杉(shan)幼枝扦(qian)(qian)插成活率較高;1978年(nian)繼續插條(tiao)繁殖,同(tong)時開始進(jin)行以(yi)(yi)日(ri)(ri)本冷杉(shan)為砧木的(de)異(yi)砧嫁接(jie)試驗,吳嗚(wu)翔、吳森茂采用(yong)圍土(tu)(tu)劈接(jie),成活三株。方法是(shi)將日(ri)(ri)本冷杉(shan)苗多(duo)蘗條(tiao)剪掉,僅留一(yi)蘗繼續生長,以(yi)(yi)吸收和制造養分,選(xuan)一(yi)較粗(cu)的(de)蘗條(tiao)剪去后作接(jie)砧,近地徑處(chu)劈接(jie),然后回(hui)復泥(ni)土(tu)(tu),以(yi)(yi)利保濕(shi),故稱圍土(tu)(tu)劈接(jie)。
1979年,吳(wu)嗚翔與(yu)吳(wu)敦序(xu)仍以圍土(tu)劈接(jie)(jie)法異砧嫁(jia)接(jie)(jie),成活(huo)2株(zhu),其中一(yi)株(zhu)接(jie)(jie)穗(sui)第(di)二年才萌芽生長,留蘗(bo)第(di)三(san)年自行枯死,可能(neng)是插穗(sui)自行發(fa)根。
1982年吳善興試驗以日(ri)本冷杉為砧(zhen)木(mu)(mu)(mu)(mu)的髓心形成層貼接(jie)(jie)(jie)法(fa)嫁(jia)(jia)接(jie)(jie)(jie),此(ci)(ci)時(shi)砧(zhen)木(mu)(mu)(mu)(mu)已長大,成活率高,長勢好;接(jie)(jie)(jie)著石柏林、沈明(ming)溫、馬海泉(quan)繼續進(jin)(jin)(jin)行(xing)(xing)貼接(jie)(jie)(jie)繁殖,共保存50佘株。此(ci)(ci)方法(fa)可應用于生產推廠,但(dan)嫁(jia)(jia)接(jie)(jie)(jie)接(jie)(jie)(jie)苗受砧(zhen)木(mu)(mu)(mu)(mu)蒙導(dao)影響,是復合體,后代(dai)是雜交種,不(bu)是純種。吳鳴翔等(deng)也在百山祖(zu)主峰進(jin)(jin)(jin)行(xing)(xing)以日(ri)本冷杉為砧(zhen)木(mu)(mu)(mu)(mu)的靠接(jie)(jie)(jie)插(cha)(cha),即插(cha)(cha)穗稈插(cha)(cha)入(ru)土,同時(shi)中部靠接(jie)(jie)(jie)于砧(zhen)木(mu)(mu)(mu)(mu)上,獲一株成活,并開過(guo)雄花;還進(jin)(jin)(jin)行(xing)(xing)過(guo)馬尾松(song),黃山松(song)異砧(zhen)嫁(jia)(jia)接(jie)(jie)(jie),沒有成活。
1986年以來,沈明溫(wen)、馬海泉相繼進(jin)行(xing)扦插試驗,試用幾種(zhong)生長素及ABT生根(gen)粉處(chu)理插穗,其中(zhong)包括(kuo)大(da)多數采用嫁接(jie)株枝條(tiao)為插穗,保存(cun)率略有提高。科(ke)研工作者相繼進(jin)行(xing)常(chang)規無(wu)性(xing)繁(fan)殖(zhi),由于百山祖冷杉僅適(shi)應中(zhong)山氣候(hou),而且無(wu)法連(lian)續(xu)剪(jian)取大(da)量繁(fan)殖(zhi)材料,易產生愈傷組織,但(dan)發根(gen)難(nan),因(yin)而成(cheng)效不大(da)。
栽培技術
為了拯救(jiu)百山祖冷(leng)杉,免于物種滅絕,必須采取綜(zong)合技術措(cuo)施,應包括二(er)個方(fang)面(mian),一是(shi)搞清稀有原因,主要是(shi)阻礙(ai)繁殖的機(ji)理,二(er)是(shi)研究繁殖技術,具體內容是(shi):
1、建立定位觀測站(zhan),搞(gao)清本(ben)群落中(zhong)植物種同競爭對百山祖冷(leng)(leng)杉(shan)的影響(xiang),包括鄰近(jin)地(di)區冷(leng)(leng)杉(shan)林(lin)群落學(xue)研(yan)究。以采用調整(zheng)種間關系的技術。
2、氣候因子對(dui)百山祖(zu)冷杉開花(hua)結實的(de)影響。降雨、低溫阻礙結實的(de)對(dui)策(ce)措施(shi)。
3、土壤(rang)(rang)因子對百山祖冷杉結實(shi)的影響,主要是搞(gao)清土壤(rang)(rang)養分的種(zhong)類,分布現狀及(ji)冷杉根系分布惰況,探(tan)討相(xiang)應的人工施肥技(ji)術。
4、百山祖冷杉生殖生理學的研究,如性別分(fen)離(li)、不育(yu)性、結(jie)實生理等,施用赤霉(mei)素促進開(kai)花,人(ren)工授(shou)粉(fen)技術。
5、嫁接繁殖:
①在雌株(zhu)上嫁接有(you)雄花的枝條;
②以(yi)日本冷杉為砧木的靠接插,成(cheng)活后立即分開,以(yi)減少砧木對(dui)接穗(sui)的影(ying)響(xiang)。
百山祖冷杉全株
百山祖冷杉全株
6、生(sheng)物(wu)工程新技術的應用:
①莖尖培(pei)養,國外已獲成(cheng)功(gong),可(ke)以應(ying)用;
②種胚培養;
③花藥培養;
④根尖(jian)或愈(yu)傷組織培養。
建議組織(zhi)專家(jia)進行論(lun)證(zheng),由百(bai)山(shan)祖(zu)自然(ran)保護區及(ji)有(you)(you)關(guan)科研(yan)所(suo)及(ji)院校(xiao)組成攻關(guan)小組,實(shi)施(shi)拯教(jiao)工作(zuo),也可爭取海外或國際(ji)援助,使(shi)舉(ju)世矚目的百(bai)山(shan)祖(zu)冷杉(shan)物(wu)種(zhong)(zhong)繁衍,欣欣向榮,快復其在(zai)舞林生(sheng)態系統中的應有(you)(you)地(di)位,在(zai)人(ren)與生(sheng)物(wu)圈中發揮應有(you)(you)的作(zuo)用(yong),實(shi)現美國國家(jia)植(zhi)物(wu)園研(yan)究(jiu)敢(gan)授戴(dai)德理博士(shi)所(suo)希(xi)望的那樣,使(shi)百(bai)山(shan)祖(zu)冷杉(shan)像水杉(shan)一樣在(zai)世界各地(di)植(zhi)物(wu)園種(zhong)(zhong)植(zhi)。
植物保護
1976年(nian)1月1日(ri),吳(wu)鳴翔陪同中(zhong)國(guo)地質(zhi)科(ke)學(xue)院的專家(jia)上(shang)山考察。一(yi)行人在沒過(guo)膝蓋的積雪中(zhong)整(zheng)整(zheng)徒步(bu)走了(le)大半天,才見到了(le)冷(leng)杉。1976年(nian)冷(leng)杉屬(shu)新種———百山祖冷(leng)杉在《植物(wu)分(fen)類學(xue)報》上(shang)正式發表(biao),引起(qi)了(le)國(guo)內外植物(wu)學(xue)界的關(guan)注。
瀕危原因
第(di)一,由(you)于全(quan)(quan)球氣候變(bian)遷(qian),逼使冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)分(fen)(fen)布區(qu)的(de)變(bian)遷(qian)。在(zai)植(zhi)物(wu)分(fen)(fen)類(lei)系統中(zhong),冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)是古(gu)老的(de)裸(luo)子植(zhi)物(wu)松(song)科中(zhong)的(de)一個屬(shu)(shu),冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)屬(shu)(shu)家族全(quan)(quan)世界(jie)有(you)50多(duo)(duo)種(zhong),按照鄭(zheng)萬鈞裸(luo)子植(zhi)物(wu)分(fen)(fen)類(lei)系統,已知(zhi)中(zhong)國(guo)原產的(de)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)屬(shu)(shu)植(zhi)物(wu)有(you)23種(zhong)及(ji)數變(bian)種(zhong),百(bai)山(shan)(shan)(shan)祖冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)是在(zai)中(zhong)國(guo)發(fa)(fa)現(xian)(xian)的(de)第(di)19種(zhong)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)。在(zai)地(di)球生物(wu)圈生物(wu)地(di)帶(dai)性群(qun)落中(zhong),冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)屬(shu)(shu)植(zhi)物(wu)多(duo)(duo)分(fen)(fen)布于寒冷(leng)(leng)(leng)(leng)(leng)(leng)濕潤的(de)高緯度(du)高海(hai)拔地(di)帶(dai)。第(di)四紀冰川時(shi)期(qi),全(quan)(quan)球氣溫下降,冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)的(de)分(fen)(fen)布區(qu)向(xiang)低緯度(du),中(zhong)低海(hai)拔擴散,江蘇(su)丹陽,江西吉安、浙江天(tian)目山(shan)(shan)(shan)(900m海(hai)拔)及(ji)長江口外現(xian)(xian)代已成(cheng)海(hai)洋的(de)"東海(hai)平原"地(di)層中(zhong)均有(you)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)孢粉發(fa)(fa)現(xian)(xian)。冰期(qi)過(guo)后(hou)全(quan)(quan)球氣溫回升,冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)不(bu)能適應高溫環(huan)境,其(qi)分(fen)(fen)布區(qu)向(xiang)高緯度(du)和高中(zhong)海(hai)撥山(shan)(shan)(shan)地(di)退縮,形成(cheng)現(xian)(xian)代的(de)我國(guo)南(nan)方冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)的(de)孤島狀分(fen)(fen)布:除原已知(zhi)臺灣(wan)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)和巴山(shan)(shan)(shan)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)分(fen)(fen)布緯度(du)較南(nan),但都在(zai)2300米以上山(shan)(shan)(shan)地(di);百(bai)山(shan)(shan)(shan)祖冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)發(fa)(fa)現(xian)(xian)后(hou),梵(fan)凈山(shan)(shan)(shan)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan),湖南(nan)發(fa)(fa)現(xian)(xian)大院(yuan)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)等新(xin)種(zhong)以及(ji)井崗山(shan)(shan)(shan)發(fa)(fa)現(xian)(xian)巴山(shan)(shan)(shan)冷(leng)(leng)(leng)(leng)(leng)(leng)杉(shan)(shan)(shan)(shan)新(xin)分(fen)(fen)布。
第二(er),由(you)于(yu)人類開發活動及(ji)森林(lin)火(huo)災,使冷(leng)杉分(fen)布區(qu)的(de)(de)"孤島(dao)"面(mian)積更為縮小,如(ru)浙西(xi)南(nan)閩東(dong)山(shan)地,歷史上(shang)由(you)于(yu)交(jiao)通阻(zu)塞(sai),森林(lin)的(de)(de)直接經(jing)濟效益無法實現,加上(shang)燒(shao)荒驅獸,形成(cheng)了一些山(shan)區(qu)盲目(mu)燒(shao)荒的(de)(de)傳統習慣,燒(shao)毀當時認為"多(duo)余的(de)(de)"森林(lin)。百山(shan)祖冷(leng)杉自然生長的(de)(de)林(lin)分(fen),由(you)于(yu)低凹穩風(feng),被燒(shao)的(de)(de)次(ci)數(shu)少得多(duo),是(shi)劫難(nan)后(hou)的(de)(de)幸存者,如(ru)被稱為二(er)號樹(shu)的(de)(de)這(zhe)株冷(leng)杉,是(shi)雪壓(ya)且火(huo)燒(shao)后(hou)萌芽(ya)長成(cheng)的(de)(de),基部還有原先樹(shu)桿(gan)的(de)(de)斷面(mian)保存著。浙南(nan)這(zhe)種類型較為原始的(de)(de)植被都(dou)是(shi)分(fen)散分(fen)布在各個山(shan)頭。
第(di)三(san),種(zhong)之間(jian)生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)存(cun)競爭激烈(lie)。森林群落的(de)主體(ti)是(shi)由(you)多(duo)(duo)(duo)(duo)種(zhong)木本植(zhi)物(wu)(wu)組(zu)成(cheng)的(de),樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)種(zhong)之間(jian)相(xiang)生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)相(xiang)克現象(xiang),在歷史(shi)的(de)長(chang)(chang)河(he)中,是(shi)隨(sui)著環境(主要是(shi)氣候)因(yin)子(zi)的(de)變(bian)(bian)遷(qian),按照(zhao)事物(wu)(wu)運(yun)動的(de)矛盾(dun)對立統(tong)一規律而(er)(er)演(yan)變(bian)(bian)的(de),是(shi)生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)物(wu)(wu)區(qu)(qu)系演(yan)變(bian)(bian)的(de)一部分(fen)。環境的(de)變(bian)(bian)化逼(bi)使生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)物(wu)(wu)適應(ying)新(xin)的(de)環境,使形態或(huo)適應(ying)能力產生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)變(bian)(bian)異分(fen)化,適者(zhe)存(cun),成(cheng)為(wei)新(xin)的(de)類群,不適者(zhe)淘汰或(huo)在環境改(gai)變(bian)(bian)不大(da)的(de)區(qu)(qu)域保存(cun)下來;同時生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)物(wu)(wu)群體(ti)組(zu)成(cheng)的(de)生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)態系統(tong)改(gai)變(bian)(bian)的(de)環境,如森林的(de)涵養水源(yuan),保持(chi)水土(tu)、防風固(gu)沙、吸(xi)收(shou)二氧化碳而(er)(er)阻緩"溫(wen)室效應(ying)"的(de)進(jin)程等(deng)(deng)。現存(cun)百(bai)山祖(zu)冷(leng)杉(shan)自然生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)長(chang)(chang)的(de)這個林分(fen),就是(shi)上述生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)物(wu)(wu)區(qu)(qu)系演(yan)變(bian)(bian)的(de)一個縮影,建群種(zhong)為(wei)亮葉(xie)(xie)水青岡和百(bai)山祖(zu)冷(leng)杉(shan),第(di)二層(ceng)為(wei)多(duo)(duo)(duo)(duo)脈(mo)青岡和紫莖等(deng)(deng);灌木層(ceng)為(wei)粉紅短柱茶及兩種(zhong)小(xiao)竹(百(bai)山祖(zu)玉山竹和慶(qing)元箬竹)等(deng)(deng);草(cao)本層(ceng)有竹節人(ren)參、開(kai)口箭和蕨類等(deng)(deng);層(ceng)外(wai)植(zhi)物(wu)(wu)有常青藤(teng)衛予、剌果(guo)毒漆(qi)藤(teng)等(deng)(deng);苔蘚植(zhi)物(wu)(wu)既有短角(jiao)管苔(原知分(fen)布歐洲(zhou),此(ci)處為(wei)洲(zhou)際新(xin)分(fen)布記錄),又有能治心紋痛的(de)暖地大(da)葉(xie)(xie)蘚,后者(zhe)顧名(ming)思義(yi)是(shi)能分(fen)布于溫(wen)暖地帶的(de)檀物(wu)(wu)。喬木層(ceng)主要樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)種(zhong)亮葉(xie)(xie)水青岡與百(bai)山祖(zu)冷(leng)杉(shan)競爭激烈(lie),冷(leng)杉(shan)被擠在體(ti)緣,在林中的(de)原3號樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)冷(leng)杉(shan),胸徑20厘米,而(er)(er)樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)冠(guan)冠(guan)幅不到(dao)1米,采(cai)取(qu)人(ren)工干(gan)預,擴(kuo)大(da)了林冠(guan)空間(jian),但(dan)仍在1986年(nian)因(yin)冷(leng)杉(shan)樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)干(gan)在亮葉(xie)(xie)水青岡枝(zhi)叉間(jian),風吹樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)杈磨損(sun)了冷(leng)杉(shan)樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)干(gan)一圈(quan)樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)皮(pi)而(er)(er)枯死。現存(cun)最小(xiao)的(de)-株冷(leng)杉(shan)1963年(nian)已采(cai)過標本,由(you)于伴生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)種(zhong)林冠(guan)閉(bi)鎖覆蓋,30多(duo)(duo)(duo)(duo)年(nian)生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)仍還高不過2米人(ren)工培土(tu)和減少庇萌(meng),才開(kai)始恢(hui)復生(sheng)(sheng)(sheng)(sheng)(sheng)(sheng)機。現存(cun)2號樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)的(de)大(da)枝(zhi)都(dou)向林緣方(fang)向屈曲(qu),以取(qu)得更多(duo)(duo)(duo)(duo)的(de)陽光,但(dan)這株樹(shu)(shu)(shu)(shu)(shu)(shu)(shu)(shu)的(de)大(da)枝(zhi)也(ye)快速枯損(sun),長(chang)(chang)勢在迅速衰退中。
第四,由于(yu)多(duo)雨的(de)(de)小氣(qi)候(hou)(hou),土壤(rang)淋浴作用強(qiang)烈,遣成土壤(rang)貧瘠。百山(shan)祖冷杉積聚養分(fen)以開(kai)花結實的(de)(de)間隔期長,開(kai)花的(de)(de)機(ji)會難得,1982年開(kai)花后到1991年才開(kai)花。根據德(de)國生態學家H.沃爾侍(Hei nri ch Walter)的(de)(de)地(di)(di)球陸地(di)(di)生物(wu)圈的(de)(de)地(di)(di)帶(dai)(dai)生物(wu)群落分(fen)類劃分(fen),浙江百山(shan)祖位于(yu)Ⅱ-v地(di)(di)帶(dai)(dai)群落交(jiao)錯區,即有夏雨的(de)(de)熱(re)帶(dai)(dai)與暖溫帶(dai)(dai)(海洋性)群落交(jiao)錯區,氣(qi)候(hou)(hou)帶(dai)(dai)為亞熱(re)帶(dai)(dai)濕潤季鳳區,夏秋(qiu)多(duo)地(di)(di)形雨,而(er)且住往花期多(duo)雨,松類是風媒花,雨水將(jiang)花粉沖洗下地(di)(di)而(er)無(wu)法飛(fei)揚,嚴(yan)重影(ying)響授(shou)粉,人(ren)工嫁接繁殖的(de)(de)植株同樣出現(xian)這(zhe)一情況(kuang)。
第五,百山(shan)(shan)祖冷(leng)(leng)杉(shan)(shan)(shan)種群內部(bu)變異(yi)分(fen)化,形成性(xing)別分(fen)離和個體(ti)生態差異(yi),也是(shi)自(zi)(zi)然(ran)有性(xing)繁(fan)殖的(de)障礙之(zhi)一。據觀察,僅(jin)存(cun)四株(zhu)(zhu)(zhu)(zhu)(zhu)冷(leng)(leng)杉(shan)(shan)(shan)中有三(san)(san)株(zhu)(zhu)(zhu)(zhu)(zhu)能開花(hua)(hua),這三(san)(san)株(zhu)(zhu)(zhu)(zhu)(zhu)中1號(hao)(hao)樹(shu)(shu)牢兩性(xing)同(tong)株(zhu)(zhu)(zhu)(zhu)(zhu),2號(hao)(hao)樹(shu)(shu)為雌性(xing)株(zhu)(zhu)(zhu)(zhu)(zhu),另一株(zhu)(zhu)(zhu)(zhu)(zhu)可(ke)能也是(shi)雌株(zhu)(zhu)(zhu)(zhu)(zhu),且2號(hao)(hao)樹(shu)(shu)遠離1號(hao)(hao)樹(shu)(shu),2號(hao)(hao)樹(shu)(shu)花(hua)(hua)期(qi)較1號(hao)(hao)樹(shu)(shu)早幾(ji)天(tian)(tian)(2號(hao)(hao)樹(shu)(shu)生長立地比1號(hao)(hao)樹(shu)(shu)海拔低);同(tong)時(shi),這幾(ji)株(zhu)(zhu)(zhu)(zhu)(zhu)樹(shu)(shu)并非同(tong)年(nian)開花(hua)(hua),1991年(nian)2號(hao)(hao)樹(shu)(shu)就未開花(hua)(hua)。第六,百山(shan)(shan)祖冷(leng)(leng)杉(shan)(shan)(shan)種群個體(ti)太少,1969年(nian)時(shi)有8株(zhu)(zhu)(zhu)(zhu)(zhu)樹(shu)(shu)。三(san)(san)株(zhu)(zhu)(zhu)(zhu)(zhu)單株(zhu)(zhu)(zhu)(zhu)(zhu)散生;另5株(zhu)(zhu)(zhu)(zhu)(zhu)組成一個小群體(ti),二(er)大三(san)(san)小,天(tian)(tian)然(ran)更新(xin)尚好,但夏天(tian)(tian)最(zui)大的(de)那株(zhu)(zhu)(zhu)(zhu)(zhu)冷(leng)(leng)杉(shan)(shan)(shan)被洪水沖倒枯死,1977年(nian)被挖2株(zhu)(zhu)(zhu)(zhu)(zhu),剩下(xia)一大一小,近(jin)親繁(fan)殖,是(shi)否(fou)影繁(fan)殖機制尚不得(de)而知。總之(zhi),百山(shan)(shan)祖冷(leng)(leng)杉(shan)(shan)(shan)瀕(bin)危稀有的(de)原(yuan)因有環境因素,也有物種本身內在因素,若仍任其(qi)自(zi)(zi)然(ran)演變,很難避免物種滅絕趨勢發展。
保護措施
1976年定(ding)名發表的百山祖冷杉,是(shi)中國浙江省百山祖自(zi)(zi)然保(bao)護區的特有(you)(you)植物,在浙江第二(er)高(gao)峰百山祖主(zhu)峰西(xi)南(nan)側1700米以(yi)上(shang)山谷溝旁的亮葉水青(qing)?林中,由于(yu)種(zhong)種(zhong)原(yuan)因,這(zhe)種(zhong)冷杉自(zi)(zi)然有(you)(you)性(xing)繁殖十分困(kun)難,常(chang)規人工(gong)無(wu)性(xing)繁殖也困(kun)難,瀕(bin)臨物種(zhong)滅絕境地。
為(wei)了引起(qi)人們的重(zhong)視,1987年2月,國際物(wu)(wu)種(zhong)保護委員(yuan)會(SSC)將百山祖冷杉公(gong)布(bu)列(lie)為(wei)世界最瀕(bin)危的十二種(zhong)植物(wu)(wu)之一(yi)。
為了搶救百(bai)山(shan)祖冷(leng)杉(shan),1991年林業(ye)(ye)專(zhuan)家(jia)采(cai)集一批百(bai)山(shan)祖冷(leng)杉(shan)種(zhong)子,1992年進行人工育苗。2000年3月,林業(ye)(ye)專(zhuan)家(jia)把(ba)這批實生苗移(yi)植到(dao)慶(qing)元縣(xian)五大堡鄉種(zhong)植,經過專(zhuan)家(jia)們10多年的精心(xin)管護(hu),遷地(di)保(bao)護(hu)的11株(zhu)百(bai)山(shan)祖冷(leng)杉(shan)實生苗株(zhu)高(gao)已達1.95米(mi),冠(guan)幅(fu)1.3米(mi),地(di)徑(jing)0.5米(mi),長勢喜人。
主要價值
冷(leng)杉(shan)是裸子植(zhi)物(wu)(wu)中(zhong)(zhong)(zhong)的一個(ge)小家(jia)族(zu),但(dan)家(jia)族(zu)成員也不少,僅中(zhong)(zhong)(zhong)國就有20多種(zhong),其中(zhong)(zhong)(zhong)7種(zhong)被列(lie)為國家(jia)保護植(zhi)物(wu)(wu)。百(bai)山祖冷(leng)杉(shan)是冷(leng)杉(shan)家(jia)族(zu)中(zhong)(zhong)(zhong)最珍(zhen)貴(gui)的種(zhong)類,有“植(zhi)物(wu)(wu)活(huo)化(hua)石”及“植(zhi)物(wu)(wu)大(da)熊貓”的美(mei)稱,已被列(lie)為國家(jia)一級保護植(zhi)物(wu)(wu)。
百山(shan)祖冷(leng)杉(shan)是(shi)中國特有的古(gu)老殘遺(yi)植物(wu),是(shi)蘇、浙、皖、閩等省唯一(yi)生(sheng)存至今的冷(leng)杉(shan)屬中的珍稀(xi)物(wu)種,對研究植物(wu)區系和氣候變遷等方面有較重要的學術意義。