桃花(hua)水(shui)(shui)母(mu)的(de)生(sheng)活史為無(wu)(wu)性生(sheng)殖與(yu)有(you)性生(sheng)殖世代交替而(er)成,螅(xi)狀(zhuang)體(ti)(水(shui)(shui)母(mu)的(de)“童年”)對(dui)環(huan)(huan)境(jing)要求極低,而(er)一(yi)旦分離出(chu)水(shui)(shui)母(mu)則對(dui)環(huan)(huan)境(jing)和(he)水(shui)(shui)質(zhi)要求很高(gao)。環(huan)(huan)境(jing)適應時,螅(xi)狀(zhuang)體(ti)便自然分離出(chu)水(shui)(shui)母(mu);環(huan)(huan)境(jing)不利時,螅(xi)狀(zhuang)體(ti)便長期吸附(fu)于(yu)水(shui)(shui)下或巖石縫(feng)中世代生(sheng)存下去。桃花(hua)水(shui)(shui)母(mu)產于(yu)淡水(shui)(shui)中,居(ju)住人工湖(hu),池(chi)塘、湖(hu)泊和(he)河(he)流。對(dui)水(shui)(shui)環(huan)(huan)境(jing)的(de)要求極高(gao),適宜其生(sheng)存的(de)水(shui)(shui)域(yu)必須無(wu)(wu)毒無(wu)(wu)害、潔凈,且(qie)多(duo)呈(cheng)酸性。測(ce)出(chu)PH值為6.4。這(zhe)一(yi)微(wei)酸水(shui)(shui)環(huan)(huan)境(jing)最(zui)適宜水(shui)(shui)母(mu)生(sheng)長。
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桃花水(shui)(shui)(shui)母(mu)(mu)直徑(jing)約15-25毫米,幾乎是透(tou)明的(de)(de),有(you)(you)(you)白色(se)(se)或(huo)綠色(se)(se)色(se)(se)調,觸角約20毫米長。呈現出傘(san)狀,有(you)(you)(you)螺紋串狀的(de)(de)觸手圍繞其圓(yuan)邊(環形緣)。圓(yuan)形管的(de)(de)直徑(jing)從5至25毫米。是僅有(you)(you)(you)的(de)(de)一種淡水(shui)(shui)(shui)生(sheng)活的(de)(de)小型(xing)(xing)(xing)(xing)水(shui)(shui)(shui)母(mu)(mu),生(sheng)活史(shi)具(ju)有(you)(you)(you)單體(ti)(ti)的(de)(de)小型(xing)(xing)(xing)(xing)的(de)(de)水(shui)(shui)(shui)螅(xi)(xi)型(xing)(xing)(xing)(xing)體(ti)(ti)和自(zi)由(you)游泳的(de)(de)水(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)體(ti)(ti),但以(yi)水(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)為主。水(shui)(shui)(shui)螅(xi)(xi)型(xing)(xing)(xing)(xing)體(ti)(ti)無圍鞘,甚至無觸手,單體(ti)(ti)常僅數(shu)毫米。水(shui)(shui)(shui)母(mu)(mu)體(ti)(ti)也(ye)為小型(xing)(xing)(xing)(xing),都具(ju)緣膜(mo),這是水(shui)(shui)(shui)螅(xi)(xi)水(shui)(shui)(shui)母(mu)(mu)的(de)(de)特征,具(ju)很多觸手。水(shui)(shui)(shui)螅(xi)(xi)型(xing)(xing)(xing)(xing)結構簡單,沒(mei)有(you)(you)(you)口道,胃(wei)循環腔中(zhong)也(ye)沒(mei)有(you)(you)(you)隔膜(mo)。水(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)絕大多數(shu)具(ju)有(you)(you)(you)緣膜(mo),胃(wei)腔中(zhong)沒(mei)有(you)(you)(you)刺(ci)細(xi)胞。水(shui)(shui)(shui)螅(xi)(xi)型(xing)(xing)(xing)(xing)及水(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)的(de)(de)中(zhong)膠層中(zhong)均無細(xi)胞結構,生(sheng)殖細(xi)胞均來(lai)源于表皮層(外胚層),個別種即使來(lai)源于內胚層,但最(zui)后仍在外胚層中(zhong)發育成熟。
桃(tao)花(hua)水(shui)(shui)(shui)母,又稱“桃(tao)花(hua)魚”、“降落傘(san)魚”,生長于溫帶(dai)淡水(shui)(shui)(shui)中(zhong),其(qi)形(xing)狀(zhuang)如桃(tao)花(hua),并(bing)多在桃(tao)花(hua)季節出現,故(gu)得名。其(qi)通體(ti)透明,像透明小傘(san)在水(shui)(shui)(shui)中(zhong)悠然漂浮,它們(men)無(wu)頭(tou)無(wu)尾呈(cheng)圓形(xing),晶瑩透亮,柔軟如綢,身體(ti)周邊長滿了(le)觸角,中(zhong)間長著(zhu)五個呈(cheng)桃(tao)花(hua)形(xing)分布的觸角狀(zhuang)物體(ti)。它們(men)在水(shui)(shui)(shui)中(zhong)一張一縮上下(xia)飄蕩,悠然自得。是一種瀕臨絕跡、古老而珍(zhen)稀的腔腸動物。
桃花(hua)水母(mu)的(de)(de)(de)(de)外(wai)形象一(yi)把撐開的(de)(de)(de)(de)傘(san),或一(yi)個倒扣的(de)(de)(de)(de)碗,稱為傘(san)體,傘(san)邊緣(yuan)有許多細線狀(zhuang)的(de)(de)(de)(de)觸(chu)手,這是它的(de)(de)(de)(de)捕食(shi)器官。口(kou)朝向傘(san)下(xia)方,位(wei)于一(yi)條管(guan)子的(de)(de)(de)(de)末(mo)端,具4片唇。食(shi)物由(you)此(ci)吞入(ru),消化不了的(de)(de)(de)(de)殘渣,仍由(you)口(kou)排出體外(wai),所以桃花(hua)水母(mu)嘴和肛門是一(yi)個。傘(san)緣(yuan)有一(yi)水平緣(yuan)膜,此(ci)為水螅(xi)水母(mu)的(de)(de)(de)(de)共同(tong)特征。
桃花水母的(de)生殖腺顏色(se)(se)有粉紅色(se)(se)、褐黃色(se)(se)、微(wei)綠色(se)(se)、淡(dan)(dan)黃色(se)(se)、綠色(se)(se)、白色(se)(se)及淡(dan)(dan)墨色(se)(se)等。均隨其種類(或種群)和變種而不同。
桃花(hua)水母(mu)體(ti)的(de)(de)傘(san)(san)體(ti)不停地收縮(suo)與舒張,將下傘(san)(san)腔(qiang)內(nei)的(de)(de)水壓出(chu)體(ti)外,借此(ci)朝相反的(de)(de)方向緩慢游動。當遇到食物時(shi),觸手上(shang)的(de)(de)刺絲囊即射出(chu)刺絲,刺中(zhong)被捕(bu)獲物,頃(qing)刻將其(qi)中(zhong)毒身亡(wang),以觸手送入口(kou)中(zhong),吞入胃內(nei)。桃花(hua)水母(mu)多以劍(jian)水蚤、小線蟲、小環蟲、小蝌蚪、小魚苗等為食。
桃花(hua)(hua)水(shui)母(mu)常出(chu)(chu)現(xian)(xian)(xian)在(zai)(zai)溫(wen)暖的(de)水(shui)域中(zhong)(zhong),溫(wen)度至(zhi)少為(wei)25℃,但水(shui)溫(wen)不能高于(yu)32℃。在(zai)(zai)初(chu)夏5月水(shui)溫(wen)14℃和(he)深秋(qiu)(qiu)11月水(shui)溫(wen)12℃時(shi)都(dou)曾采到,其生存(cun)水(shui)域的(de)pH為(wei)6.5-8.0,以每年6-10月的(de)夏秋(qiu)(qiu)季節出(chu)(chu)現(xian)(xian)(xian)最多(duo)。這是因為(wei)適宜的(de)水(shui)溫(wen)和(he)充足的(de)食物使其大量繁(fan)殖,往(wang)(wang)往(wang)(wang)在(zai)(zai)7-9月達(da)到高峰,10月底或(huo)(huo)11月初(chu)后桃花(hua)(hua)水(shui)母(mu)群體數量迅速(su)下降(jiang),此后便銷聲匿跡。同一地點常年(或(huo)(huo)每年)出(chu)(chu)現(xian)(xian)(xian)的(de)現(xian)(xian)(xian)象十分罕見。桃花(hua)(hua)水(shui)母(mu)對日光(guang)有正(zheng)向反應,喜弱光(guang),有一定(ding)的(de)趨(qu)光(guang)性,早、晚(wan)分布(bu)在(zai)(zai)水(shui)的(de)上層(ceng)或(huo)(huo)表層(ceng),中(zhong)(zhong)午光(guang)照強、水(shui)溫(wen)高時(shi)則多(duo)分布(bu)于(yu)水(shui)的(de)中(zhong)(zhong)下層(ceng)。故在(zai)(zai)晴朗無(wu)風的(de)早晨和(he)傍晚(wan)時(shi)更(geng)易采到。
每到暮春,桃花(hua)怒放時(shi),它們(men)(men)就相繼出現。桃花(hua)水(shui)(shui)母有(you)粉紅色(se)的生殖(zhi)腺,透明的傘頂(ding),在水(shui)(shui)中浮沉漂蕩,有(you)如落水(shui)(shui)桃花(hua),因(yin)此,中國古代(dai)稱(cheng)它們(men)(men)為(wei)“桃花(hua)魚(yu)”。但又明確指出,桃花(hua)魚(yu)“非魚(yu)也”,因(yin)其水(shui)(shui)生,故名之曰魚(yu);又因(yin)其生于桃花(hua)開時(shi),故名之曰“桃花(hua)魚(yu)”。
自然界(jie)的(de)(de)桃花(hua)(hua)水(shui)(shui)(shui)母(mu)在(zai)低溫、富營(ying)養(yang)(yang)及饑餓的(de)(de)不利環(huan)境條(tiao)(tiao)件(jian)下,螅狀體(ti)(ti)可(ke)收縮,分泌(mi)一層堅硬的(de)(de)保護膜包被(bei)(bei)身體(ti)(ti),成為休(xiu)眠體(ti)(ti)(podocyst)。有學者(zhe)(zhe)認(ren)為,休(xiu)眠體(ti)(ti)能(neng)通過水(shui)(shui)(shui)生(sheng)植物(wu)、動物(wu)(如(ru)鳥(niao))或(huo)人為的(de)(de)因素,而使其(qi)傳(chuan)播。環(huan)境條(tiao)(tiao)件(jian)改善時,休(xiu)眠體(ti)(ti)重新發育為螅狀體(ti)(ti),其(qi)生(sheng)活(huo)史得以(yi)繼續。所(suo)以(yi)桃花(hua)(hua)水(shui)(shui)(shui)母(mu)也(ye)能(neng)在(zai)房頂(ding)水(shui)(shui)(shui)箱中被(bei)(bei)發現。也(ye)有研究者(zhe)(zhe)認(ren)為,桃花(hua)(hua)水(shui)(shui)(shui)母(mu)喜生(sheng)活(huo)在(zai)富營(ying)養(yang)(yang)的(de)(de)池塘,且勝過寡營(ying)養(yang)(yang)的(de)(de)大湖(hu)泊,寡營(ying)養(yang)(yang)的(de)(de)水(shui)(shui)(shui)域中缺少浮(fu)游植物(wu),當然也(ye)影響小型浮(fu)游動物(wu)的(de)(de)繁殖,桃花(hua)(hua)水(shui)(shui)(shui)母(mu)就(jiu)沒(mei)有了(le)生(sheng)存(cun)的(de)(de)餌料條(tiao)(tiao)件(jian),只要不是長期(qi)的(de)(de)嚴重的(de)(de)富營(ying)養(yang)(yang)化的(de)(de)水(shui)(shui)(shui)域,還(huan)是有可(ke)能(neng)出現桃花(hua)(hua)水(shui)(shui)(shui)母(mu)的(de)(de)。
桃花水(shui)母在何種水(shui)體中出現(xian)有(you)兩種原因決定:其(qi)(qi)一,它對(dui)水(shui)質要求極高,周(zhou)邊環境(jing)的(de)細微變化(hua)都會導致其(qi)(qi)消失;其(qi)(qi)二,桃花水(shui)母無(wu)性(xing)生殖世(shi)代的(de)水(shui)螅型(xing)不(bu)發達(da),螅狀體大小僅有(you)0.5-1.0mm,附著(zhu)在水(shui)下腐敗植物、巖石和(he)樹樁等物體上,體表(biao)有(you)粘(zhan)液,常粘(zhan)附著(zhu)一些碎屑(xie),將身體部分地遮蓋,一般極難被(bei)發現(xian)和(he)認識,其(qi)(qi)萌發成水(shui)母體的(de)機制尚不(bu)清楚。
桃花水(shui)(shui)母屬(shu)水(shui)(shui)母型(xing)生殖體(ti)(ti)。個體(ti)(ti)呈(cheng)退化(hua)的水(shui)(shui)母型(xing)結構(gou),無口(kou)、無觸手及感官,具有大量的生殖細胞(bao),雌(ci)雄(xiong)異(yi)體(ti)(ti),但群體(ti)(ti)可(ke)以是(shi)雌(ci)雄(xiong)同體(ti)(ti),生殖細胞(bao)在釋(shi)放之后個體(ti)(ti)死去。刺(ci)胞(bao)動(dong)(dong)物(wu)(wu)中(zhong)(zhong)多態(tai)現(xian)象的生物(wu)(wu)學意義(yi)可(ke)以認(ren)為是(shi)群體(ti)(ti)中(zhong)(zhong)個體(ti)(ti)之間的勞動(dong)(dong)分工,因為腔腸(chang)(chang)動(dong)(dong)物(wu)(wu)尚(shang)未出(chu)現(xian)器官系統來(lai)擔任不同的生理機(ji)能(neng),而(er)是(shi)通過(guo)群體(ti)(ti)中(zhong)(zhong)個體(ti)(ti)的形(xing)態(tai)分化(hua)來(lai)擔任不同的生理機(ji)能(neng),這(zhe)種分化(hua)是(shi)一種很原(yuan)始的形(xing)式,這(zhe)與腔腸(chang)(chang)動(dong)(dong)物(wu)(wu)的生活史及發展水(shui)(shui)平(ping)是(shi)密切相關的。
身體呈水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing),生(sheng)(sheng)活(huo)史很簡單,即由水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)體產(chan)生(sheng)(sheng)精(jing)、卵(luan),并形(xing)成(cheng)受精(jing)卵(luan)→浮(fu)浪幼蟲(chong)→水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)單體。生(sheng)(sheng)活(huo)史中沒有(you)(you)水(shui)(shui)(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing),有(you)(you)的種(zhong)甚至沒有(you)(you)浮(fu)浪幼蟲(chong)期(qi)。在二(er)態(tai)及多(duo)態(tai)生(sheng)(sheng)活(huo)的種(zhong)類,生(sheng)(sheng)活(huo)史中有(you)(you)水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)及水(shui)(shui)(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)階段,其水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)行無性生(sheng)(sheng)殖,水(shui)(shui)(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)或(huo)水(shui)(shui)(shui)(shui)(shui)母(mu)(mu)芽行有(you)(you)性生(sheng)(sheng)殖,其受精(jing)卵(luan)仍發育成(cheng)水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)體,其生(sheng)(sheng)活(huo)史成(cheng)為水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)體→水(shui)(shui)(shui)(shui)(shui)母(mu)(mu)型(xing)(xing)(xing)(xing)體或(huo)水(shui)(shui)(shui)(shui)(shui)母(mu)(mu)芽→受精(jing)卵(luan)→浮(fu)浪幼蟲(chong)→水(shui)(shui)(shui)(shui)(shui)螅(xi)型(xing)(xing)(xing)(xing)體。結果形(xing)成(cheng)無性世代與有(you)(you)性世代的交(jiao)替(ti)現象(xiang)。
按照前一種(zhong)觀(guan)(guan)點(dian),原始的(de)腔(qiang)腸動(dong)物(wu)(wu)應(ying)(ying)是(shi)(shi)水(shui)(shui)螅型(xing)(xing)(xing)(xing),它通(tong)過有性(xing)生殖,次生性(xing)的(de)產生水(shui)(shui)母(mu)型(xing)(xing)(xing)(xing)體(ti),因(yin)此(ci)形成(cheng)(cheng)(cheng)了二態或多態。按照后(hou)(hou)一種(zhong)觀(guan)(guan)點(dian)認為(wei)(wei),祖先(xian)(xian)腔(qiang)腸動(dong)物(wu)(wu)是(shi)(shi)柔軟的(de)水(shui)(shui)母(mu)型(xing)(xing)(xing)(xing),而水(shui)(shui)螅型(xing)(xing)(xing)(xing)是(shi)(shi)幼(you)蟲(chong)(chong)期的(de)持續。這后(hou)(hou)一種(zhong)觀(guan)(guan)點(dian)在(zai)水(shui)(shui)螅綱硬(ying)(ying)水(shui)(shui)母(mu)目中得到了證實。因(yin)為(wei)(wei)硬(ying)(ying)水(shui)(shui)母(mu)類(lei)的(de)受精卵在(zai)發(fa)(fa)育(yu)中通(tong)過浮(fu)(fu)(fu)浪(lang)幼(you)蟲(chong)(chong)及輻(fu)射(she)幼(you)蟲(chong)(chong)(actinula)直接發(fa)(fa)育(yu)成(cheng)(cheng)(cheng)水(shui)(shui)母(mu)型(xing)(xing)(xing)(xing),其生活(huo)史(shi)是(shi)(shi)水(shui)(shui)母(mu)型(xing)(xing)(xing)(xing)體(ti)→受精卵→浮(fu)(fu)(fu)浪(lang)幼(you)蟲(chong)(chong)→輻(fu)射(she)幼(you)蟲(chong)(chong)→水(shui)(shui)母(mu)體(ti)。一種(zhong)小型(xing)(xing)(xing)(xing)的(de)水(shui)(shui)螅Microhydra的(de)個體(ti)發(fa)(fa)育(yu)正(zheng)是(shi)(shi)如此(ci),這說明輻(fu)射(she)幼(you)蟲(chong)(chong)首先(xian)(xian)發(fa)(fa)展了固著(zhu)習性(xing),隨著(zhu)延(yan)伸(shen)固著(zhu)期,停(ting)止發(fa)(fa)育(yu)成(cheng)(cheng)(cheng)水(shui)(shui)母(mu)型(xing)(xing)(xing)(xing),而最(zui)終發(fa)(fa)育(yu)成(cheng)(cheng)(cheng)水(shui)(shui)螅型(xing)(xing)(xing)(xing)群(qun)體(ti),所以生活(huo)史(shi)的(de)改變(bian)形成(cheng)(cheng)(cheng)了多態。這種(zhong)現(xian)象與管水(shui)(shui)母(mu)類(lei)的(de)發(fa)(fa)育(yu)也(ye)相符合,它的(de)浮(fu)(fu)(fu)浪(lang)幼(you)蟲(chong)(chong)也(ye)是(shi)(shi)首先(xian)(xian)形成(cheng)(cheng)(cheng)水(shui)(shui)母(mu)型(xing)(xing)(xing)(xing)體(ti),即浮(fu)(fu)(fu)囊體(ti)、游泳體(ti)。因(yin)此(ci)世代交替的(de)觀(guan)(guan)點(dian)應(ying)(ying)該(gai)讓位于水(shui)(shui)母(mu)體(ti)是(shi)(shi)成(cheng)(cheng)(cheng)熟的(de)刺胞動(dong)物(wu)(wu),而水(shui)(shui)螅體(ti)是(shi)(shi)幼(you)蟲(chong)(chong)期的(de)持續階段(duan)的(de)觀(guan)(guan)點(dian)。當然這個論(lun)點(dian)是(shi)(shi)建筑在(zai)硬(ying)(ying)水(shui)(shui)母(mu)目是(shi)(shi)最(zui)原始的(de)刺胞動(dong)物(wu)(wu)的(de)基(ji)礎之上(shang)的(de)。
桃花水(shui)(shui)(shui)母(mu)(mu)(mu)突然出(chu)(chu)現,經數(shu)日或十數(shu)日后又悄(qiao)悄(qiao)地消失,這要從它們的(de)(de)生殖規律來解答(da)。中國在(zai)早春(chun)看到的(de)(de)桃花水(shui)(shui)(shui)母(mu)(mu)(mu),為(wei)成熟的(de)(de)水(shui)(shui)(shui)母(mu)(mu)(mu)體(ti)(ti)(ti),有(you)雌(ci)雄之分。觸手(shou)細長,伸向上(shang)方的(de)(de)均為(wei)雌(ci)性(xing)(xing),觸手(shou)短粗,垂向下方的(de)(de)為(wei)雄性(xing)(xing)。雌(ci)雄將卵(luan)子和(he)精子排到水(shui)(shui)(shui)中,卵(luan)受精發(fa)(fa)育成一個微小的(de)(de)滿被纖毛的(de)(de)浮游幼蟲(chong),后幼蟲(chong)一端接觸石(shi)土央等外(wai)物,固著(zhu),發(fa)(fa)育成一個極小的(de)(de)樹枝(zhi)狀的(de)(de)水(shui)(shui)(shui)螅(xi)(xi)體(ti)(ti)(ti)。水(shui)(shui)(shui)螅(xi)(xi)體(ti)(ti)(ti)可(ke)度(du)過酷熱的(de)(de)夏季(ji)和(he)嚴寒(han)的(de)(de)冬季(ji),待來年春(chun)天,以出(chu)(chu)芽生殖產(chan)生水(shui)(shui)(shui)母(mu)(mu)(mu)體(ti)(ti)(ti)。水(shui)(shui)(shui)母(mu)(mu)(mu)體(ti)(ti)(ti)成熟后,再進行有(you)性(xing)(xing)生殖。水(shui)(shui)(shui)螅(xi)(xi)體(ti)(ti)(ti)長0.3毫米,水(shui)(shui)(shui)母(mu)(mu)(mu)體(ti)(ti)(ti)進行有(you)性(xing)(xing)生殖產(chan)生水(shui)(shui)(shui)螅(xi)(xi)體(ti)(ti)(ti),水(shui)(shui)(shui)螅(xi)(xi)體(ti)(ti)(ti)進行無性(xing)(xing)生殖產(chan)生水(shui)(shui)(shui)母(mu)(mu)(mu)體(ti)(ti)(ti),這在(zai)動物學上(shang)稱為(wei)世(shi)代交替。