瓢(piao)(piao)(piao)蟲(chong)(chong)繭(jian)蜂Perilitus coccinellae(Schrank)是(shi)繭(jian)蜂科優(you)繭(jian)蜂亞(ya)科緣繭(jian)蜂屬的(de)一(yi)個物種。瓢(piao)(piao)(piao)蟲(chong)(chong)繭(jian)蜂幼(you)蟲(chong)(chong)操控瓢(piao)(piao)(piao)蟲(chong)(chong)保護(hu)蜂繭(jian)的(de)時間(jian)長度存在(zai)差異。有(you)時候瓢(piao)(piao)(piao)蟲(chong)(chong)會一(yi)直保持警惕(ti),直到幼(you)蟲(chong)(chong)破繭(jian)而出,蛻變成(cheng)黃蜂。以(yi)幼(you)蟲(chong)(chong)在(zai)瓢(piao)(piao)(piao)蟲(chong)(chong)體內和一(yi)部分以(yi)繭(jian)在(zai)田(tian)間(jian)越冬,春季3-4月羽化為(wei)成(cheng)蟲(chong)(chong)。寄(ji)主有(you)七星瓤蟲(chong)(chong)、稻紅瓢(piao)(piao)(piao)蟲(chong)(chong)。成(cheng)蟲(chong)(chong)寄(ji)生,單寄(ji)生。
北京(海淀(dian),延慶,懷柔),山(shan)西(xi)(花塔,榆次(ci),臨汾),內蒙古(賀蘭山(shan)),浙江(西(xi)天目山(shan),東陽),福(fu)建(jian)(福(fu)州,漳州,龍溪),河南(nan)(安(an)陽),四川(chuan)(西(xi)昌(chang)),云南(nan)(昆明,開(kai)遠(yuan)),新(xin)疆(烏魯木齊)。
全世界(包括夏威(wei)夷和新(xin)西蘭)。
雌蜂(feng)體長約4毫米。頭頂黃褐(he)色(se),顏面(mian)黃色(se),復眼、單(dan)眼座(zuo)及(ji)后頭黑(hei)色(se),觸角由(you)基(ji)部(bu)至端部(bu)深(shen)褐(he)色(se)至黑(hei)色(se);胸(xiong)部(bu)及(ji)并胸(xiong)腹(fu)(fu)節黑(hei)色(se);足暗(an)褐(he)色(se),前足稍淡;翅(chi)透(tou)明,略帶煙褐(he)色(se),翅(chi)痣與(yu)翅(chi)脈暗(an)褐(he)色(se);腹(fu)(fu)部(bu),第1背(bei)板端部(bu)變(bian)寬具刻紋(wen)及(ji)皺(zhou)狀(zhuang)刻點,第2節及(ji)其后各節背(bei)板平滑,具光澤。產卵(luan)管突出,直(zhi),產卵(luan)管鞘長約為腹(fu)(fu)長之半(ban),被均(jun)勻(yun)白毛(mao)。
頭(tou)光(guang)滑(hua)具(ju)白色(se)柔毛(mao)(mao),顏面中(zhong)央(yang)略(lve)隆起,與(yu)唇基(ji)間(jian)分界明顯,唇基(ji)窩深,有(you)后(hou)(hou)頭(tou)脊;觸(chu)角與(yu)體約等長,22節。中(zhong)胸(xiong)盾片(pian)中(zhong)葉(xie)(xie)前半部(bu)及(ji)側葉(xie)(xie)后(hou)(hou)半部(bu)光(guang)滑(hua)具(ju)光(guang)澤,稍(shao)隆起,盾縱溝寬而淺,有(you)網狀(zhuang)皺(zhou)紋,至盾片(pian)中(zhong)葉(xie)(xie)前半部(bu)連成一(yi)片(pian),似“U”字形(xing);小盾片(pian)具(ju)網紋;并胸(xiong)腹(fu)(fu)節具(ju)粗網紋,自基(ji)部(bu)1/3以后(hou)(hou)向(xiang)下傾(qing)斜,后(hou)(hou)半部(bu)有(you)細中(zhong)脊。前翅徑(jing)室上(shang)緣短(duan)于徑(jing)脈(mo)(mo)端部(bu)距(ju)翅端的距(ju)離(li),徑(jing)脈(mo)(mo)第1段(duan)很短(duan),約與(yu)基(ji)徑(jing)基(ji)段(duan)等長,無(wu)第2肘間(jian)脈(mo)(mo),回脈(mo)(mo)自肘室伸(shen)出,小脈(mo)(mo)后(hou)(hou)叉式(shi)。腹(fu)(fu)部(bu)第1節有(you)柄,具(ju)網狀(zhuang)皺(zhou)紋,第2節以后(hou)(hou)各節背板光(guang)滑(hua),具(ju)光(guang)澤。產卵(luan)管(guan)鞘黑(hei)色(se),約為(wei)腹(fu)(fu)長之半,被有(you)白色(se)細毛(mao)(mao)。幼(you)蟲(chong)長橢圓形(xing),黃白毛(mao)(mao),頭(tou)部(bu)較粗,后(hou)(hou)端較細,無(wu)足,腹(fu)(fu)末(mo)有(you)水泡狀(zhuang)突起。老熟時體長約5.5毫(hao)(hao)米(mi)(mi)。繭長約5毫(hao)(hao)米(mi)(mi),紡(fang)錘形(xing),黃褐色(se),繭外(wai)有(you)一(yi)層(ceng)疏松的深灰黃色(se)薄絲。單繭。
寄(ji)(ji)主有七星瓤(rang)蟲(chong)(chong)、稻(dao)(dao)紅瓢蟲(chong)(chong)。成蟲(chong)(chong)寄(ji)(ji)生(sheng),單寄(ji)(ji)生(sheng)。蜂幼(you)蟲(chong)(chong)老熟時鉆出(chu)(chu)寄(ji)(ji)主體外(wai),在其腹下吐絲結繭(jian)(jian),繭(jian)(jian)白色,并有一(yi)(yi)(yi)絲與瓤(rang)蟲(chong)(chong)相連,此時蟲(chong)(chong)已進人垂死(si)狀態,俯在蜂繭(jian)(jian)之(zhi)上,尤(you)如(ru)孵蛋(dan)一(yi)(yi)(yi)樣,觸之(zhi)能微動(dong)(dong),不食不動(dong)(dong)維持十天左右,寄(ji)(ji)生(sheng)蜂才咬破繭(jian)(jian)的(de)一(yi)(yi)(yi)端(切口整齊)羽(yu)化出(chu)(chu)來。貴定地區四月底在麥田可(ke)見七星瓢蟲(chong)(chong)被寄(ji)(ji)生(sheng);9月間(jian)可(ke)見稻(dao)(dao)田稻(dao)(dao)紅艦蟲(chong)(chong)被寄(ji)(ji)生(sheng)。另發現有一(yi)(yi)(yi)種金小蜂作重(zhong)寄(ji)(ji)生(sheng),從蟲(chong)(chong)繭(jian)(jian)蜂繭(jian)(jian)內羽(yu)化出(chu)(chu)來。
黃蜂(feng)家族的瓢(piao)蟲繭(jian)蜂(feng)將(jiang)瓢(piao)蟲變(bian)成(cheng)(cheng)僵尸的全過程。瓢(piao)蟲繭(jian)蜂(feng)鎖定目標,準備(bei)將(jiang)一(yi)(yi)枚卵注入瓢(piao)蟲體(ti)內。在(zai)蜂(feng)毒(du)的侵襲下,瓢(piao)蟲將(jiang)陷(xian)入癱瘓狀態。在(zai)瓢(piao)蟲體(ti)內,卵孵(fu)化成(cheng)(cheng)幼蟲,發(fa)育幾(ji)天(tian)之(zhi)后(hou)便在(zai)瓢(piao)蟲腹部抓出(chu)一(yi)(yi)個(ge)小(xiao)洞(dong)。隨(sui)后(hou),幼蟲在(zai)瓢(piao)蟲腿(tui)之(zhi)間(jian)織出(chu)一(yi)(yi)個(ge)繭(jian),幼蟲蛻變(bian)期間(jian),瓢(piao)蟲的尸體(ti)一(yi)(yi)直安息在(zai)繭(jian)頂(ding)部。
研(yan)究(jiu)指(zhi)出絕(jue)大多數寄生蟲(chong)(chong)最終都殺死宿主(zhu),一些(xie)瓢(piao)蟲(chong)(chong)經(jing)受住可(ke)怕(pa)的考驗,幸(xing)存(cun)下(xia)來。30%到40%的瓢(piao)蟲(chong)(chong)宿主(zhu)在黃(huang)蜂幼蟲(chong)(chong)孵(fu)化之(zhi)后仍然存(cun)活,其中一些(xie)個體還能(neng)繼續產卵。這(zhe)些(xie)瓢(piao)蟲(chong)(chong)之(zhi)所以能(neng)夠(gou)幸(xing)存(cun)的原(yuan)因在于,黃(huang)蜂幼蟲(chong)(chong)只(zhi)以并不對瓢(piao)蟲(chong)(chong)構成致命威脅的組織為(wei)食,例如脂肪。
仍(reng)然活(huo)著(zhu)的(de)瓢(piao)蟲(chong)(chong)站在蜂(feng)繭上面(mian),此時的(de)瓢(piao)蟲(chong)(chong)具有(you)攻擊(ji)性(xing)(xing),攻擊(ji)入侵(qin)者(zhe)。這種行為與(yu)黃蜂(feng)幼蟲(chong)(chong)鉆出并結繭后(hou)留在瓢(piao)蟲(chong)(chong)體(ti)內的(de)毒(du)液(ye)有(you)關。研(yan)究小組將捕食(shi)性(xing)(xing)草蜻蛉(ling)放入培養(yang)皿(min),蜂(feng)繭有(you)的(de)上面(mian)站著(zhu)活(huo)瓢(piao)蟲(chong)(chong),有(you)的(de)站著(zhu)死蜂(feng)繭,有(you)的(de)則沒有(you)這個保(bao)鏢(biao)。實驗結果顯示,草蜻蛉(ling)在攻擊(ji)受到僵尸(shi)保(bao)鏢(biao)保(bao)護的(de)蜂(feng)繭時成功率較低。
瓢(piao)蟲(chong)(chong)(chong)繭蜂幼蟲(chong)(chong)(chong)操控(kong)(kong)瓢(piao)蟲(chong)(chong)(chong)保(bao)(bao)護(hu)蜂繭的時(shi)間(jian)長度存在差異。有(you)時(shi)候(hou)瓢(piao)蟲(chong)(chong)(chong)會一(yi)(yi)直保(bao)(bao)持警惕(ti),直到幼蟲(chong)(chong)(chong)破(po)繭而出,蛻變成黃(huang)蜂。其(qi)他情況下,瓢(piao)蟲(chong)(chong)(chong)被控(kong)(kong)制的時(shi)間(jian)只有(you)幾天。研究發現,如果投入更多時(shi)間(jian)和能量控(kong)(kong)制它們的瓢(piao)蟲(chong)(chong)(chong)保(bao)(bao)鏢,黃(huang)蜂幼蟲(chong)(chong)(chong)成年后產卵(luan)較少。這(zhe)是(shi)科學家第一(yi)(yi)次發現控(kong)(kong)制宿主(zhu)與自身繁殖(zhi)之(zhi)間(jian)的取(qu)舍(she)關系。
最終,一(yi)只新黃蜂(feng)從(cong)受到保護(hu)的繭中鉆出。布(bu)羅德表示(shi)這(zhe)項研究(jiu)揭(jie)開了加拿(na)大的一(yi)個(ge)謎團。他說:“在森林中散步時,我(wo)(wo)們(men)經常看(kan)到瓢蟲站在一(yi)個(ge)繭上面,這(zhe)讓(rang)我(wo)(wo)們(men)大為迷惑。現在,我(wo)(wo)們(men)終于找到答案。”
以(yi)幼(you)(you)蟲(chong)(chong)(chong)在(zai)(zai)(zai)瓢蟲(chong)(chong)(chong)體(ti)內(nei)和一部分(fen)以(yi)繭在(zai)(zai)(zai)田間(jian)越冬,春季(ji)3-4月(yue)(yue)(yue)羽(yu)化為成蟲(chong)(chong)(chong),產卵(luan)于(yu)瓢蟲(chong)(chong)(chong)腹(fu)末或腹(fu)側的節間(jian)膜內(nei),單寄生,幼(you)(you)蟲(chong)(chong)(chong)老熟(shu)后(hou)在(zai)(zai)(zai)寄主腹(fu)部背面第(di)4-5節間(jian)咬圓孔(kong)外出(chu),在(zai)(zai)(zai)瓢蟲(chong)(chong)(chong)體(ti)下(xia)吐(tu)絲作繭。成蟲(chong)(chong)(chong)壽命平(ping)均(jun)(jun)7天,蛹期(qi)平(ping)均(jun)(jun)14天,卵(luan)和幼(you)(you)蟲(chong)(chong)(chong)期(qi)共43天,第(di)1代(dai)幼(you)(you)蟲(chong)(chong)(chong)期(qi)在(zai)(zai)(zai)4月(yue)(yue)(yue)上旬至4月(yue)(yue)(yue)底(di),第(di)2代(dai)在(zai)(zai)(zai)6月(yue)(yue)(yue)。
生境:農田(tian)、果園、森林。
地理分布(bu):河北、山東、河南(nan)、陜(shan)西、上海、四川、臺灣(wan)、福(fu)建、廣東、廣西、云南(nan)。