南(nan)(nan)(nan)方巨(ju)獸(shou)龍(long)(long)(long)(long)屬下(xia)僅(jin)有(you)一(yi)(yi)種,為(wei)卡氏南(nan)(nan)(nan)方巨(ju)獸(shou)龍(long)(long)(long)(long),是(shi)一(yi)(yi)種發現于阿根廷的(de)大(da)型(xing)獸(shou)腳類恐(kong)龍(long)(long)(long)(long),生存年(nian)(nian)代(dai)為(wei)晚白堊世塞諾(nuo)曼期,約距今9700萬年(nian)(nian)前。南(nan)(nan)(nan)方巨(ju)獸(shou)龍(long)(long)(long)(long)屬于異特(te)龍(long)(long)(long)(long)超科中的(de)鯊齒龍(long)(long)(long)(long)類,并(bing)且很有(you)可能是(shi)地(di)球歷史中體(ti)型(xing)最大(da)的(de)陸地(di)掠食動物。與霸王(wang)龍(long)(long)(long)(long)相(xiang)(xiang)比(bi),南(nan)(nan)(nan)方巨(ju)獸(shou)的(de)樣本(ben)量(liang)很小,且大(da)個體(ti)標本(ben)的(de)完整(zheng)度(du)很差,因(yin)此對其體(ti)型(xing)的(de)準確(que)估計有(you)相(xiang)(xiang)當的(de)難度(du)。它與霸王(wang)龍(long)(long)(long)(long)相(xiang)(xiang)比(bi)誰的(de)體(ti)型(xing)更大(da)也是(shi)學術界一(yi)(yi)直在爭論(lun)的(de)話題。
南(nan)方巨獸龍(long)屬名的含義是巨型(xing)南(nan)方蜥(xi)蜴,這是由于它發現(xian)于南(nan)方大陸且體型(xing)巨大。種名卡氏是為了紀(ji)念化石的發現(xian)者卡洛琳(Rubén D.Carolini)。
南方巨(ju)獸龍的(de)正(zheng)型(xing)標本(ben)(ben)(ben)約(yue)有70%的(de)完整度,包(bao)含(han)了大多數(shu)的(de)脊椎骨(gu),肩帶(dai)和腰帶(dai)骨(gu)骼,股骨(gu),以(yi)及左側(ce)的(de)脛骨(gu)和腓骨(gu)。這件(jian)標本(ben)(ben)(ben)于1995年(nian)被描述,相(xiang)關成果發表在自然科學頂級(ji)雜志(zhi)Nature上。1998年(nian),科里亞等人(ren)(ren)又將帶(dai)有少數(shu)牙齒(chi)的(de)一(yi)(yi)塊下頜(he)歸(gui)入到(dao)南方巨(ju)獸龍。這件(jian)下頜(he)標本(ben)(ben)(ben)比1995年(nian)描述的(de)正(zheng)型(xing)標本(ben)(ben)(ben)還(huan)要(yao)大一(yi)(yi)些,科里亞等人(ren)(ren)甚至(zhi)認為這件(jian)標本(ben)(ben)(ben)的(de)頭骨(gu)總長達到(dao)1.95米。
南(nan)(nan)方(fang)巨(ju)獸(shou)(shou)(shou)(shou)龍(long)的(de)(de)(de)(de)(de)頭(tou)骨(gu)(gu)(gu)(gu)整(zheng)體較(jiao)為(wei)(wei)低矮,其(qi)上(shang)(shang)(shang)(shang)頜骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)齒(chi)(chi)列(lie)長度(du)約為(wei)(wei)92厘米(mi)。其(qi)上(shang)(shang)(shang)(shang)頜骨(gu)(gu)(gu)(gu)具(ju)(ju)有(you)一(yi)(yi)個(ge)(ge)(ge)(ge)(ge)非常(chang)發(fa)達(da)的(de)(de)(de)(de)(de)鼻(bi)骨(gu)(gu)(gu)(gu)下(xia)突(tu)和(he)(he)一(yi)(yi)個(ge)(ge)(ge)(ge)(ge)較(jiao)小(xiao)的(de)(de)(de)(de)(de)橢(tuo)圓形上(shang)(shang)(shang)(shang)頜孔。鼻(bi)骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)背面非常(chang)粗(cu)糙,充滿褶皺(zhou)。這樣(yang)粗(cu)糙的(de)(de)(de)(de)(de)表面向(xiang)(xiang)頭(tou)骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)后向(xiang)(xiang)延申,并(bing)覆(fu)蓋(gai)整(zheng)個(ge)(ge)(ge)(ge)(ge)鼻(bi)骨(gu)(gu)(gu)(gu)背面。淚骨(gu)(gu)(gu)(gu)在前突(tu)和(he)(he)下(xia)突(tu)交匯的(de)(de)(de)(de)(de)地方(fang)形成(cheng)一(yi)(yi)個(ge)(ge)(ge)(ge)(ge)非常(chang)發(fa)達(da)的(de)(de)(de)(de)(de)脊,這一(yi)(yi)點與很(hen)(hen)多異特龍(long)類相似(si)。眶后骨(gu)(gu)(gu)(gu)下(xia)突(tu)與顴(quan)骨(gu)(gu)(gu)(gu)相接處具(ju)(ju)備一(yi)(yi)向(xiang)(xiang)眼眶內深入的(de)(de)(de)(de)(de)突(tu)起。方(fang)骨(gu)(gu)(gu)(gu)約44厘米(mi)長,其(qi)內部(bu)具(ju)(ju)有(you)氣腔化的(de)(de)(de)(de)(de)結構。額骨(gu)(gu)(gu)(gu)和(he)(he)頂骨(gu)(gu)(gu)(gu)寬闊(kuo),對上(shang)(shang)(shang)(shang)顳窩(wo)造成(cheng)了(le)一(yi)(yi)定程(cheng)度(du)的(de)(de)(de)(de)(de)遮擋。齒(chi)(chi)骨(gu)(gu)(gu)(gu)在齒(chi)(chi)骨(gu)(gu)(gu)(gu)聯合(he)處縱向(xiang)(xiang)加高(gao)。每一(yi)(yi)個(ge)(ge)(ge)(ge)(ge)齒(chi)(chi)骨(gu)(gu)(gu)(gu)可能有(you)12個(ge)(ge)(ge)(ge)(ge)齒(chi)(chi)窩(wo)。除了(le)第(di)一(yi)(yi)顆牙(ya)齒(chi)(chi)比較(jiao)小(xiao)以(yi)(yi)外,下(xia)頜齒(chi)(chi)的(de)(de)(de)(de)(de)大小(xiao)都(dou)(dou)比較(jiao)接近。牙(ya)齒(chi)(chi)比較(jiao)薄(bo),在前緣和(he)(he)后緣處都(dou)(dou)具(ju)(ju)有(you)鋸齒(chi)(chi),鋸齒(chi)(chi)密度(du)約為(wei)(wei)一(yi)(yi)毫米(mi)9-12個(ge)(ge)(ge)(ge)(ge)鋸齒(chi)(chi)。牙(ya)齒(chi)(chi)的(de)(de)(de)(de)(de)表面具(ju)(ju)有(you)明顯(xian)的(de)(de)(de)(de)(de)釉質褶皺(zhou)。生物(wu)力學研(yan)究發(fa)現,南(nan)(nan)方(fang)巨(ju)獸(shou)(shou)(shou)(shou)龍(long)的(de)(de)(de)(de)(de)咬合(he)力弱于霸(ba)王(wang)龍(long)。因此,古生物(wu)學家認為(wei)(wei)南(nan)(nan)方(fang)巨(ju)獸(shou)(shou)(shou)(shou)龍(long)的(de)(de)(de)(de)(de)大嘴(zui)更適合(he)撕開獵物(wu)的(de)(de)(de)(de)(de)皮肉,而非霸(ba)王(wang)龍(long)那(nei)樣(yang)直接壓(ya)碎獵物(wu)的(de)(de)(de)(de)(de)骨(gu)(gu)(gu)(gu)骼。南(nan)(nan)方(fang)巨(ju)獸(shou)(shou)(shou)(shou)龍(long)很(hen)(hen)有(you)可能以(yi)(yi)同時代的(de)(de)(de)(de)(de)泰坦巨(ju)龍(long)類為(wei)(wei)食。
南方巨獸龍的(de)(de)頸(jing)部非(fei)常強壯(zhuang)(zhuang),具(ju)有(you)(you)粗(cu)壯(zhuang)(zhuang)的(de)(de)樞椎(zhui)。其余的(de)(de)頸(jing)椎(zhui)具(ju)有(you)(you)短且略扁的(de)(de)椎(zhui)體,椎(zhui)體內部有(you)(you)明顯的(de)(de)氣腔(qiang)化構(gou)造。尾(wei)椎(zhui)的(de)(de)椎(zhui)體粗(cu)壯(zhuang)(zhuang),前端(duan)尾(wei)椎(zhui)的(de)(de)脈弓呈刀片狀(zhuang)。肩(jian)帶骨(gu)(gu)(gu)骼(ge)較(jiao)短,肩(jian)胛骨(gu)(gu)(gu)的(de)(de)長(chang)度(du)(du)少(shao)于股骨(gu)(gu)(gu)長(chang)度(du)(du)的(de)(de)一半。肩(jian)胛骨(gu)(gu)(gu)柄具(ju)有(you)(you)很強壯(zhuang)(zhuang)的(de)(de)肌肉附著點。腸骨(gu)(gu)(gu)的(de)(de)北緣(yuan)向(xiang)上(shang)拱起,髖(kuan)臼(jiu)窩后突低(di)矮(ai),尾(wei)骨(gu)(gu)(gu)短肌附著脊較(jiao)窄。恥骨(gu)(gu)(gu)遠端(duan)的(de)(de)突起發達,且前緣(yuan)較(jiao)后緣(yuan)短。股骨(gu)(gu)(gu)呈S形。股骨(gu)(gu)(gu)頭非(fei)常強壯(zhuang)(zhuang),并指(zhi)向(xiang)內上(shang)側(ce)。小轉子(zi)呈翅膀狀(zhuang),其近端(duan)位(wei)于大轉子(zi)之下。第四轉子(zi)發達并指(zhi)向(xiang)后方。
目(mu)前的(de)(de)主流(liu)觀點認為南(nan)(nan)(nan)方(fang)巨(ju)(ju)獸龍(long)(long)(long)屬于異特龍(long)(long)(long)超科的(de)(de)鯊(sha)齒龍(long)(long)(long)類,與同屬于這一類群的(de)(de)馬普龍(long)(long)(long)構成姐(jie)妹(mei)單元。這一類群中不乏巨(ju)(ju)型化的(de)(de)捕食者(zhe),例如非洲的(de)(de)鯊(sha)齒龍(long)(long)(long),以及同樣發現于南(nan)(nan)(nan)美的(de)(de)魁(kui)紂龍(long)(long)(long)和馬普龍(long)(long)(long),還(huan)有(you)北美洲的(de)(de)高棘龍(long)(long)(long)和西(xi)雅茨龍(long)(long)(long),它們(men)的(de)(de)體(ti)型都與霸王龍(long)(long)(long)十分接近甚至更為巨(ju)(ju)大。這些巨(ju)(ju)型的(de)(de)異特龍(long)(long)(long)類統(tong)治(zhi)著早(zao)白(bai)堊世(shi)至晚白(bai)堊世(shi)初(chu)期的(de)(de)陸地(di)生態系統(tong)。在(zai)它們(men)滅絕以后,它們(men)在(zai)北半球的(de)(de)統(tong)治(zhi)權移(yi)交給了(le)暴龍(long)(long)(long)類,而在(zai)南(nan)(nan)(nan)半球的(de)(de)位置則由大型的(de)(de)阿(a)貝力龍(long)(long)(long)類取代。
南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)最(zui)大(da)(da)的(de)(de)(de)(de)(de)(de)(de)科學意(yi)義之(zhi)一(yi)(yi)就在于(yu)(yu)它可(ke)(ke)能刷新(xin)了(le)人們對(dui)于(yu)(yu)陸(lu)(lu)地(di)(di)食肉動(dong)物(wu)體(ti)(ti)(ti)型(xing)(xing)極限的(de)(de)(de)(de)(de)(de)(de)認知早期的(de)(de)(de)(de)(de)(de)(de)研究認為南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu)(gu)(gu)(gu)可(ke)(ke)能長(chang)達1.8米(mi)以上。在2001年(nian)的(de)(de)(de)(de)(de)(de)(de)一(yi)(yi)項研究中(zhong),科學家給(gei)出(chu)了(le)南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)(de)(de)(de)(de)(de)體(ti)(ti)(ti)重(zhong)估(gu)(gu)計為6.6噸(dun)。2002年(nian),科里亞(ya)與柯里在對(dui)南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)(de)(de)(de)(de)(de)腦顱進行形態描述的(de)(de)(de)(de)(de)(de)(de)時(shi)候給(gei)出(chu)了(le)南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)正型(xing)(xing)標本(ben)(ben)頭(tou)骨(gu)(gu)(gu)(gu)長(chang)度的(de)(de)(de)(de)(de)(de)(de)估(gu)(gu)計,大(da)(da)約為1.6米(mi),并依(yi)據股(gu)(gu)骨(gu)(gu)(gu)(gu)中(zhong)段的(de)(de)(de)(de)(de)(de)(de)截面的(de)(de)(de)(de)(de)(de)(de)周長(chang)估(gu)(gu)計得到體(ti)(ti)(ti)重(zhong)約為4.16噸(dun)。2004年(nian)的(de)(de)(de)(de)(de)(de)(de)一(yi)(yi)項研究通過(guo)股(gu)(gu)骨(gu)(gu)(gu)(gu)和脛腓骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)(de)(de)多(duo)種測量(liang)數據對(dui)體(ti)(ti)(ti)重(zhong)進行估(gu)(gu)算,認為南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)正型(xing)(xing)標本(ben)(ben)的(de)(de)(de)(de)(de)(de)(de)體(ti)(ti)(ti)重(zhong)可(ke)(ke)能為6.604噸(dun),而更(geng)(geng)大(da)(da)的(de)(de)(de)(de)(de)(de)(de)那具標本(ben)(ben)可(ke)(ke)能為8.47噸(dun)。如果(guo)這(zhe)一(yi)(yi)估(gu)(gu)計可(ke)(ke)靠的(de)(de)(de)(de)(de)(de)(de)話,那么(me)南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)確實是歷(li)史上最(zui)大(da)(da)的(de)(de)(de)(de)(de)(de)(de)陸(lu)(lu)地(di)(di)肉食動(dong)物(wu)。當然相(xiang)比(bi)于(yu)(yu)霸王(wang)龍(long)(long)(long)(long)(long)(long)(long)擁(yong)有(you)(you)(you)非常多(duo)的(de)(de)(de)(de)(de)(de)(de)保存(cun)精美的(de)(de)(de)(de)(de)(de)(de)標本(ben)(ben),以及(ji)較為完(wan)整(zheng)的(de)(de)(de)(de)(de)(de)(de)個體(ti)(ti)(ti)發(fa)育序列,南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)在標本(ben)(ben)數量(liang)以及(ji)標本(ben)(ben)的(de)(de)(de)(de)(de)(de)(de)完(wan)整(zheng)程度都不及(ji)前(qian)(qian)者,并且(qie)頭(tou)骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)(de)(de)上多(duo)個骨(gu)(gu)(gu)(gu)塊的(de)(de)(de)(de)(de)(de)(de)關(guan)節面沒(mei)有(you)(you)(you)保存(cun),因(yin)此(ci)使得推算頭(tou)長(chang)以及(ji)體(ti)(ti)(ti)型(xing)(xing)變得極為困(kun)難。同時(shi),鯊齒龍(long)(long)(long)(long)(long)(long)(long)類(lei)可(ke)(ke)能擁(yong)有(you)(you)(you)比(bi)暴龍(long)(long)(long)(long)(long)(long)(long)類(lei)更(geng)(geng)大(da)(da)的(de)(de)(de)(de)(de)(de)(de)頭(tou)身比(bi),因(yin)此(ci),更(geng)(geng)大(da)(da)的(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu)(gu)(gu)(gu)不一(yi)(yi)定代表南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)就擁(yong)有(you)(you)(you)比(bi)霸王(wang)龍(long)(long)(long)(long)(long)(long)(long)更(geng)(geng)大(da)(da)的(de)(de)(de)(de)(de)(de)(de)體(ti)(ti)(ti)型(xing)(xing)。另外,南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)正型(xing)(xing)標本(ben)(ben)的(de)(de)(de)(de)(de)(de)(de)股(gu)(gu)骨(gu)(gu)(gu)(gu)雖然比(bi)霸王(wang)龍(long)(long)(long)(long)(long)(long)(long)“蘇”的(de)(de)(de)(de)(de)(de)(de)股(gu)(gu)骨(gu)(gu)(gu)(gu)更(geng)(geng)強(qiang)壯更(geng)(geng)長(chang),但是它的(de)(de)(de)(de)(de)(de)(de)脛骨(gu)(gu)(gu)(gu)卻更(geng)(geng)短。短期內(nei),它和霸王(wang)龍(long)(long)(long)(long)(long)(long)(long)相(xiang)比(bi)誰大(da)(da)誰小(xiao)還很難確認。但毫無疑問(wen)的(de)(de)(de)(de)(de)(de)(de)一(yi)(yi)點(dian)是,南(nan)(nan)方(fang)(fang)巨(ju)(ju)(ju)(ju)獸(shou)(shou)(shou)(shou)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)(de)(de)(de)(de)(de)發(fa)現動(dong)搖了(le)霸王(wang)龍(long)(long)(long)(long)(long)(long)(long)王(wang)位(wei),并揭示了(le)一(yi)(yi)個在大(da)(da)型(xing)(xing)暴龍(long)(long)(long)(long)(long)(long)(long)類(lei)崛起之(zhi)前(qian)(qian)由巨(ju)(ju)(ju)(ju)型(xing)(xing)異特龍(long)(long)(long)(long)(long)(long)(long)類(lei)統治的(de)(de)(de)(de)(de)(de)(de)世界,同時(shi)對(dui)于(yu)(yu)研究在白(bai)堊紀陸(lu)(lu)地(di)(di)革命(Cretaceous Terrestrial Revolution)期間恐龍(long)(long)(long)(long)(long)(long)(long)動(dong)物(wu)類(lei)群的(de)(de)(de)(de)(de)(de)(de)轉變具有(you)(you)(you)重(zhong)要意(yi)義。
在(zai)(zai)電影侏羅(luo)紀世(shi)界(jie)3中,南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)被(bei)(bei)描(miao)述為(wei)地球歷史(shi)上最大(da)的(de)(de)陸(lu)地肉食動物,比霸(ba)(ba)(ba)王龍(long)(long)(long)(long)(long)更為(wei)巨(ju)(ju)大(da)。在(zai)(zai)電影的(de)(de)開頭,一只噩兆龍(long)(long)(long)(long)(long)(一種小型(xing)暴龍(long)(long)(long)(long)(long)類(lei)(lei),與(yu)西亞(ya)茨龍(long)(long)(long)(long)(long)發現(xian)于(yu)同(tong)一地區)在(zai)(zai)進食南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)嘴中的(de)(de)食物殘渣,之后南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)與(yu)霸(ba)(ba)(ba)王龍(long)(long)(long)(long)(long)大(da)戰(zhan)(zhan)并殺死對手(shou)。在(zai)(zai)最終的(de)(de)大(da)戰(zhan)(zhan)中,被(bei)(bei)霸(ba)(ba)(ba)王龍(long)(long)(long)(long)(long)和鐮刀(dao)龍(long)(long)(long)(long)(long)聯手(shou)殺死。導演所設(she)計的(de)(de)劇情(qing)似乎暗示了巨(ju)(ju)型(xing)異特龍(long)(long)(long)(long)(long)類(lei)(lei)與(yu)暴龍(long)(long)(long)(long)(long)類(lei)(lei)在(zai)(zai)漫長的(de)(de)演化(hua)歷史(shi)中綿(mian)延8000多萬年(nian)的(de)(de)糾(jiu)葛(ge)。然而(er),在(zai)(zai)真實的(de)(de)情(qing)況下,南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)與(yu)霸(ba)(ba)(ba)王龍(long)(long)(long)(long)(long)是(shi)不可(ke)(ke)能相遇(yu)的(de)(de),因為(wei)兩者的(de)(de)生存年(nian)代(dai)差了3000多萬年(nian)。除了侏羅(luo)紀世(shi)界(jie)3以(yi)外,南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)還出現(xian)在(zai)(zai)BBC的(de)(de)紀錄(lu)片《與(yu)龍(long)(long)(long)(long)(long)同(tong)行(xing)特輯:尋爪記》和紀錄(lu)片《巴塔哥尼(ni)亞(ya)的(de)(de)巨(ju)(ju)獸(shou)》,在(zai)(zai)這兩部紀錄(lu)片中,南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)被(bei)(bei)描(miao)述為(wei)群(qun)體獵殺阿根(gen)廷(ting)(ting)龍(long)(long)(long)(long)(long)的(de)(de)巨(ju)(ju)型(xing)獵手(shou)。阿根(gen)廷(ting)(ting)龍(long)(long)(long)(long)(long)是(shi)最大(da)的(de)(de)恐龍(long)(long)(long)(long)(long),也(ye)(ye)是(shi)最大(da)的(de)(de)陸(lu)地脊椎動物的(de)(de)種子選手(shou)之一,體長可(ke)(ke)達約(yue)40米,其化(hua)石同(tong)樣被(bei)(bei)發現(xian)于(yu)阿根(gen)廷(ting)(ting)。然而(er)南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)可(ke)(ke)能與(yu)阿根(gen)廷(ting)(ting)龍(long)(long)(long)(long)(long)不處于(yu)同(tong)一時代(dai),反而(er)是(shi)它的(de)(de)近(jin)親(qin)馬普(pu)龍(long)(long)(long)(long)(long)更可(ke)(ke)能是(shi)阿根(gen)廷(ting)(ting)龍(long)(long)(long)(long)(long)的(de)(de)天敵。南(nan)方(fang)(fang)(fang)巨(ju)(ju)獸(shou)龍(long)(long)(long)(long)(long)也(ye)(ye)出現(xian)在(zai)(zai)英劇《遠古入侵》中,被(bei)(bei)親(qin)切的(de)(de)成為(wei)G.rex。
南方巨(ju)獸(shou)龍生存于1億年(nian)前中(zhong)生代中(zhong)白堊世(shi)森諾曼期巴塔哥尼(ni)亞。
恐龍(long)種類(lei):蜥臀目(mu)·獸腳類(lei)·堅(jian)尾龍(long)類(lei).肉食龍(long)下目(mu).異特龍(long)超科.鯊(sha)齒(chi)龍(long)科.鯊(sha)齒(chi)龍(long)亞科.南方巨獸龍(long)族
1993年阿根廷一個驚人發現,南(nan)方(fang)巨獸(shou)龍正模被發現了。
1993年(nian),考古(gu)學家(jia)ruben d. carolini在(zai)1993年(nian)在(zai)阿根廷(ting)巴塔哥尼亞平原進行考古(gu)發(fa)(fa)掘的時候意外地發(fa)(fa)現(xian)了(le)一個重(zhong)大(da)的秘密(mi), 原來(lai)在(zai)遠古(gu)的阿根廷(ting)曾經存在(zai)過(guo)一種(zhong)可(ke)怕的怪獸。這種(zhong)可(ke)怕的怪獸是(shi)地球上有史以來(lai)第二大(da)的兩足生(sheng)物,最(zui)大(da)體長(chang)13米,最(zui)大(da)體重(zhong)可(ke)以達(da)到11噸,這種(zhong)恐龍(long)1995年(nian)被命名為(wei)南方巨(ju)獸龍(long),意思(si)是(shi)南方巨(ju)大(da)的蜥蜴。
MUCPv-Ch1,長12米,體(ti)重8.5噸,40%的完整(zheng)骨架,部分(fen)頭骨、一點零碎的小部分(fen)前肢(zhi)、完整(zheng)脖子、絕大(da)部分(fen)軀干、部分(fen)后肢(zhi)、大(da)部分(fen)尾巴。
MUCPv-95,長13米,體(ti)重(zhong)11噸,一個完整(zheng)的下頜骨。
南(nan)(nan)方(fang)(fang)巨(ju)獸龍(long)(long)的(de)(de)(de)(de)腦袋就(jiu)身體而言較大,下(xia)巴(ba)(ba)略(lve)呈(cheng)方(fang)(fang)形。它(ta)碩(shuo)大的(de)(de)(de)(de)嘴巴(ba)(ba)長(chang)著一(yi)口鋒(feng)利的(de)(de)(de)(de)牙齒,每顆牙有(you)9厘(li)米長(chang)。巨(ju)獸龍(long)(long)作為鯊齒龍(long)(long)科的(de)(de)(de)(de)成員(yuan),有(you)個又(you)(you)細又(you)(you)尖又(you)(you)長(chang)的(de)(de)(de)(de)尾巴(ba)(ba)。巨(ju)獸龍(long)(long)走路(lu)時(shi)用兩條腿。南(nan)(nan)方(fang)(fang)巨(ju)獸龍(long)(long)的(de)(de)(de)(de)頭骨長(chang)達(da)1.8米左(zuo)右(you),顎部長(chang)滿9-13厘(li)米長(chang)的(de)(de)(de)(de)鋒(feng)利牙齒。南(nan)(nan)方(fang)(fang)巨(ju)獸龍(long)(long)的(de)(de)(de)(de)體重(zhong)最重(zhong)可達(da)10.52噸,所以它(ta)們發展出強大的(de)(de)(de)(de)骨骼及肌肉網絡來支撐(cheng)沉重(zhong)的(de)(de)(de)(de)身軀,同時(shi)保證它(ta)們在捕食獵物的(de)(de)(de)(de)時(shi)候有(you)可接受的(de)(de)(de)(de)速(su)度。長(chang)長(chang)的(de)(de)(de)(de)尾巴(ba)(ba)則在快(kuai)(kuai)速(su)奔跑的(de)(de)(de)(de)過程中起了(le)平(ping)衡的(de)(de)(de)(de)作用和快(kuai)(kuai)速(su)轉(zhuan)向(xiang)的(de)(de)(de)(de)能力(li)。
LENGTH IN THEROPODS給出了13.1米,13.8噸(dun),但是顯然這個計算完全忽略(lve)了恐龍(long)體(ti)形的(de)差(cha)別,和頭(tou)骨的(de)體(ti)積,僅僅計算長(chang)度(du),因此可靠性非常低。
第一個南方巨(ju)獸龍和(he)霸王龍(FNMH PR 2081)的對比:
FNMH PR 2081俗名"蘇(su)",1990年出土,也是"完整腿骨的(de)霸王(wang)龍(long)"中最大的(de)(所有霸王(wang)龍(long)中不(bu)綜合年齡,蘇(su)第4/5大,綜合年齡以后并列第6)
頭骨中線(xian)長度(midline):南方(fang)巨獸龍(long)MUCPv-Ch1 1530mm,蘇1394mm
頭(tou)骨全長:南方巨獸龍MUCPv-Ch1 1700mm,蘇1435mm
下(xia)頜骨長(chang)度:南方巨獸龍MUCPv-Ch1(推(tui)測)1800mm,蘇1395mm
大腿骨長度:南方巨獸龍MUCPv-Ch1 1430mm,蘇(su)1380mm
大腿(tui)骨周長:南方(fang)巨獸龍MUCPv-Ch1 520mm,蘇580mm
小腿骨長度:南方巨獸龍MUCPv-Ch1 1120mm,蘇1200mm
頭骨寬(kuan)度:南方巨(ju)獸龍MUCPv-Ch1,520mm,蘇860mm
大小腿骨聯(lian)合長度:南方巨獸龍MUCPv-Ch1 2540mm,蘇2580mm
骨盆大小:南方巨獸(shou)龍MUCPv-Ch1 1180mm,蘇1220mm
頭骨體積(ji):霸王龍(long)略大。
2004年Mazzeta在(zai)其論文中也指出MUCPv-Ch1的(de)體(ti)(ti)積和一(yi)般的(de)霸(ba)王(wang)龍相(xiang)等,"體(ti)(ti)積"比蘇小2.5噸,但是至少和一(yi)般的(de)霸(ba)王(wang)龍體(ti)(ti)重相(xiang)等,可以從圖片看出南方巨(ju)獸龍比較瘦,和身體(ti)(ti)長(chang)度(du)接近的(de)霸(ba)王(wang)龍相(xiang)比明顯單薄一(yi)些。
蘇的(de)(de)體重(zhong)在(zai)(zai)2004年Henderson保(bao)守估(gu)算(suan)為(wei)10噸(dun),Mazzeta在(zai)(zai)估(gu)算(suan)南方巨獸龍(long)(long)(long)(long)的(de)(de)論文里面認為(wei)蘇的(de)(de)體重(zhong)8噸(dun)左右,但2013年研究蘇有13噸(dun)重(zhong)。部分資料根據(ju)保(bao)羅和(he)Henderson 1999年對(dui)一只10.7米(mi)霸王龍(long)(long)(long)(long)ANMH5027,估(gu)測上限7.9噸(dun)的(de)(de),把放大,蘇11.3噸(dun),而根據(ju)Anderson1990年對(dui)10.7米(mi)的(de)(de)雄霸王龍(long)(long)(long)(long)的(de)(de)估(gu)算(suan)是4.5噸(dun),較粗壯(zhuang)的(de)(de)雌龍(long)(long)(long)(long)蘇的(de)(de)體重(zhong)7.4噸(dun)
那(nei)么南方巨獸龍是(shi)否(fou)比一般的(de)霸王龍重(zhong)?答(da)案是(shi):比霸王龍略輕
Seebacher在(zai)2001年(nian)的(de)計算估計南(nan)方巨(ju)獸(shou)龍正(zheng)模MUCPv-Ch1 6594kg,霸王龍正(zheng)模CM 9380的(de)體(ti)重6650kg
Coria在最初公布南巨的時候給出了6000+
南方巨獸龍MUCPv-Ch1的體重的最低估測是(shi)4160kg,
南方巨獸龍的(de)體(ti)重(zhong)(zhong)進行過最詳細計(ji)算的(de)Mazzeta給出(chu)了可(ke)靠(kao)(kao)上限(xian)8.48噸,最可(ke)靠(kao)(kao)體(ti)重(zhong)(zhong)6.604-6.8噸,Mazzeta也認(ren)為MUCPv-Ch1體(ti)重(zhong)(zhong)下(xia)限(xian)4160kg是(shi)(shi)計(ji)算方式不可(ke)靠(kao)(kao),否定了南方巨獸龍4160kg的(de)說法。Mazzeta認(ren)可(ke)的(de)下(xia)限(xian)是(shi)(shi)4860kg
Mazzeta認為比MUCPv-Ch1長8%的MUCPv-95,按三次方計算,可達8.5-11.5噸。體(ti)重超過(guo)了(le)蘇,但仍舊(jiu)沒(mei)有霸(ba)王(wang)龍中MOR008,LACM 23844,MOR1126,UCMP118742重。但神(shen)秘的最大(da)南方巨獸(shou)龍超過(guo)了(le)最大(da)霸(ba)王(wang)龍MOR1126。
南方巨獸龍MUCPv-Ch1總結大小:13米,8.5噸
南(nan)方(fang)巨獸龍MUCPv-95總結大小:14米.11.5噸(dun)
其他最大獸腳類
撒哈拉鯊齒(chi)龍:13.4米,7-11.4噸
玫瑰(gui)馬(ma)普龍MCF-PVPH-108.185:14.4米,8-13.25噸
玫瑰馬(ma)普龍(long)MCF-PVPH-108.145:13.4米,7.2-11噸(dun)
玫瑰馬普龍MCF-PVPH-108.202:12.2米,7-8噸
霸王龍(long)(FNMH PR2081)12.2米,7.4-11噸(dun)
霸王龍(MOR008) 13.4米(mi),8.5-13噸(dun)
紂魁龍:12米,8.5噸(dun)
埃(ai)及棘龍MSNM 4047,17-18米
南方(fang)巨獸(shou)龍(long)是(shi)南美洲歷(li)史上(shang)最(zui)大陸(lu)地肉食(shi)動物和最(zui)大的(de)食(shi)肉恐(kong)龍(long)。也是(shi)歷(li)史上(shang)第2大的(de)食(shi)肉恐(kong)龍(long)。
南美洲中生代最大(da)的鱷魚(yu)巴(ba)西(xi)哈氏(shi)肌(ji)鱷(最大(da)個(ge)體估計哈氏(shi)帝(di)王鱷10-11米(mi),3-4噸(dun),種群體形9-10米(mi),3噸(dun)。)哈氏(shi)肌(ji)鱷是非(fei)洲帝(di)王肌(ji)鱷的近親(qin),唯一化石是一塊有11顆牙槽的43厘(li)米(mi)的下(xia)頜骨殘
片(pian),大概帝(di)王鱷對應部分是50厘米。
南方巨(ju)(ju)獸龍(long)(long)(long)出土后不久(jiu)因(yin)為(wei)體(ti)型巨(ju)(ju)大(da)(da)被認為(wei)超過霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long),因(yin)為(wei)最(zui)大(da)(da)的(de)南方巨(ju)(ju)獸龍(long)(long)(long)大(da)(da)于霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)蘇最(zui)小的(de)體(ti)重(zhong)估測。然而(er)霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)蘇的(de)體(ti)重(zhong)根據其他版本可以估測到12.8噸,遠超過南方巨(ju)(ju)獸龍(long)(long)(long)。霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)還有多個更大(da)(da)的(de)標本,可以證明霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)的(de)體(ti)重(zhong)更大(da)(da),甚至可能達到17.72噸