特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)是(shi)發(fa)現于(yu)(yu)(yu)亞洲地(di)區的(de)大(da)型暴(bao)(bao)(bao)(bao)龍(long)(long)類(lei),在外(wai)形上與北美洲的(de)霸王(wang)龍(long)(long)非(fei)(fei)常相似,是(shi)當時生態系統中(zhong)的(de)頂級捕食者。特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)體(ti)型比霸王(wang)龍(long)(long)和諸城暴(bao)(bao)(bao)(bao)龍(long)(long)略小,但(dan)明(ming)顯大(da)于(yu)(yu)(yu)其它的(de)暴(bao)(bao)(bao)(bao)龍(long)(long)類(lei),其頭骨(gu)的(de)最大(da)長度可達1.3米。特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)化(hua)石(shi)主要發(fa)現于(yu)(yu)(yu)蒙古(gu)(gu),但(dan)是(shi)也有一(yi)些證據表明(ming)這(zhe)種巨型掠食者在中(zhong)國的(de)內(nei)蒙古(gu)(gu)以及新疆等地(di)區都(dou)有分布。如同它在北美洲的(de)近親一(yi)樣(yang),特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)標(biao)本量也非(fei)(fei)常巨大(da)。截止(zhi)到目前為止(zhi),古(gu)(gu)生物學家至少已經(jing)找到了約30個特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)個體(ti)的(de)標(biao)本,這(zhe)其中(zhong)不乏保存精美的(de)頭骨(gu)化(hua)石(shi)。在亞洲恐龍(long)(long)古(gu)(gu)生物學發(fa)展(zhan)的(de)早(zao)期階段,特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)不同發(fa)育階段的(de)個體(ti)曾經(jing)被當成過(guo)不同的(de)物種,不過(guo)在近期的(de)研(yan)究中(zhong)這(zhe)些錯誤都(dou)被一(yi)一(yi)糾正。
特暴龍(long)的(de)屬名意為“可怕的(de)蜥蜴(yi)”,種名意為“勇士(shi)”。特暴龍(long)的(de)正型(xing)標(biao)本最(zui)初(chu)并(bing)沒(mei)有(you)被歸(gui)入(ru)(ru)到特暴龍(long)屬,而(er)是被歸(gui)入(ru)(ru)到暴龍(long)屬,并(bing)命名為“Tyrannosaurus bataar”,另(ling)外三個早期的(de)歸(gui)入(ru)(ru)標(biao)本中有(you)兩個被歸(gui)入(ru)(ru)了蛇發(fa)女怪龍(long)屬。直到1965年蘇聯(lian)古生物學家阿納托利·康斯坦丁諾維奇·羅(luo)特杰斯特文斯基才(cai)發(fa)現馬列夫命名的(de)這4件(jian)標(biao)本是同一物種不同發(fa)育階段的(de)個體(ti),隨后它們才(cai)被并(bing)入(ru)(ru)勇士(shi)特暴龍(long)。
正型標本(ben):PIN 551-1保存了一部分頭(tou)骨和頸椎(zhui)。最早被歸(gui)(gui)入(ru)暴龍屬,后(hou)歸(gui)(gui)入(ru)特暴龍屬。
此外(wai)蘭平暴(bao)(bao)龍(long)(Tyrannosaurus lanpingensis),吐魯(lu)番(fan)暴(bao)(bao)龍(long)(Tyrannosaurus turpanensis),Albertosaurus periculosus目前都被(bei)(bei)認為(wei)是(shi)屬于勇士特(te)暴(bao)(bao)龍(long)的(de)(de)(de)不同(tong)個(ge)體(ti)(ti)發育階段。除(chu)此之外(wai),在蒙古發現(xian)的(de)(de)(de)分支龍(long)曾經也被(bei)(bei)懷(huai)疑過是(shi)特(te)暴(bao)(bao)龍(long)的(de)(de)(de)幼年個(ge)體(ti)(ti),不過近年來(lai)由于虔州龍(long)的(de)(de)(de)發現(xian),分支龍(long)類的(de)(de)(de)有效性已經被(bei)(bei)學術(shu)界(jie)所接受。
特暴龍(long)的(de)(de)(de)頭骨(gu)(gu)側(ce)(ce)視與(yu)霸(ba)王龍(long)非常相似。兩(liang)者都具有(you)背(bei)(bei)腹向加深(shen)的(de)(de)(de)頭骨(gu)(gu),且(qie)具有(you)粗壯的(de)(de)(de)下頜。從背(bei)(bei)視可以看出兩(liang)種(zhong)恐龍(long)的(de)(de)(de)頭骨(gu)(gu)整體輪(lun)廓的(de)(de)(de)差異(yi),霸(ba)王龍(long)的(de)(de)(de)后端(duan)(duan)向外(wai)側(ce)(ce)極度(du)的(de)(de)(de)擴(kuo)展(zhan),使得雙眼視覺范(fan)圍具有(you)較大(da)的(de)(de)(de)重疊。雖然特暴龍(long)的(de)(de)(de)頭骨(gu)(gu)后端(duan)(duan)也(ye)有(you)一(yi)定(ding)程度(du)的(de)(de)(de)向外(wai)擴(kuo)展(zhan),但是(shi)與(yu)霸(ba)王龍(long)有(you)較大(da)的(de)(de)(de)差距(ju)。同時特暴龍(long)的(de)(de)(de)吻(wen)端(duan)(duan)明顯比霸(ba)王龍(long)更(geng)窄。霸(ba)王龍(long)的(de)(de)(de)前頜骨(gu)(gu)和上頜骨(gu)(gu)相接(jie)的(de)(de)(de)骨(gu)(gu)縫上有(you)一(yi)個明顯的(de)(de)(de)開孔暴露在頭骨(gu)(gu)的(de)(de)(de)外(wai)側(ce)(ce)面(mian),而這一(yi)開孔在特暴龍(long)頭骨(gu)(gu)的(de)(de)(de)外(wai)側(ce)(ce)面(mian)幾乎沒有(you)。
特(te)(te)暴龍(long)的(de)(de)(de)上(shang)頜(he)(he)骨(gu)(gu)升突粗(cu)狀發達(da),霸王(wang)龍(long)的(de)(de)(de)較(jiao)(jiao)為纖細。兩(liang)種恐龍(long)的(de)(de)(de)鼻(bi)(bi)骨(gu)(gu)腹外側緣都具(ju)有與上(shang)頜(he)(he)骨(gu)(gu)相關(guan)節的(de)(de)(de)溝,并幾乎前后向延伸至鼻(bi)(bi)骨(gu)(gu)的(de)(de)(de)整個長度。但特(te)(te)暴龍(long)的(de)(de)(de)關(guan)節面(mian)具(ju)有很多脊,使鼻(bi)(bi)骨(gu)(gu)與上(shang)頜(he)(he)骨(gu)(gu)的(de)(de)(de)關(guan)節更(geng)緊密,而(er)霸王(wang)龍(long)的(de)(de)(de)鼻(bi)(bi)骨(gu)(gu)上(shang)頜(he)(he)骨(gu)(gu)關(guan)節更(geng)簡單。相反的(de)(de)(de)是,霸王(wang)龍(long)的(de)(de)(de)鼻(bi)(bi)骨(gu)(gu)與淚骨(gu)(gu)的(de)(de)(de)關(guan)節十分復雜且非(fei)常緊密,而(er)特(te)(te)暴龍(long)的(de)(de)(de)鼻(bi)(bi)骨(gu)(gu)與淚骨(gu)(gu)的(de)(de)(de)關(guan)節則較(jiao)(jiao)為簡單。此(ci)外,霸王(wang)龍(long)的(de)(de)(de)鼻(bi)(bi)骨(gu)(gu)比(bi)特(te)(te)暴龍(long)的(de)(de)(de)更(geng)加寬大。
霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)的(de)(de)(de)(de)淚(lei)(lei)(lei)骨(gu)背緣較(jiao)特(te)暴(bao)龍(long)(long)(long)(long)有更明顯的(de)(de)(de)(de)膨大。特(te)暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)淚(lei)(lei)(lei)骨(gu)降(jiang)支(zhi)較(jiao)為中空,而霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)的(de)(de)(de)(de)淚(lei)(lei)(lei)骨(gu)降(jiang)支(zhi)幾乎是實(shi)心的(de)(de)(de)(de)。特(te)暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)淚(lei)(lei)(lei)骨(gu)前突與霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)的(de)(de)(de)(de)相(xiang)比更為細長,且(qie)前突關節面沒有分支(zhi)。霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)的(de)(de)(de)(de)淚(lei)(lei)(lei)骨(gu)前突關節面具有明顯的(de)(de)(de)(de)分叉(cha)。淚(lei)(lei)(lei)骨(gu)前突的(de)(de)(de)(de)內側面中段具有一(yi)個容納血(xue)管的(de)(de)(de)(de)開(kai)孔,這一(yi)開(kai)孔在霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)的(de)(de)(de)(de)淚(lei)(lei)(lei)骨(gu)上(shang)的(de)(de)(de)(de)位(wei)置更靠前。
鱗骨的(de)(de)(de)降支(zhi)與方(fang)顴(quan)骨的(de)(de)(de)上(shang)(shang)支(zhi)具(ju)有(you)很大(da)的(de)(de)(de)接觸面積,與艾伯塔龍(long)(long)(long)(long)(long)和蛇發女怪龍(long)(long)(long)(long)(long)不(bu)(bu)同(tong),而(er)與霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)非常相(xiang)(xiang)似。在腹(fu)(fu)側(ce)(ce)視,特暴(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)顴(quan)骨不(bu)(bu)如霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)那(nei)樣很明(ming)顯(xian)的(de)(de)(de)外翻(fan)。特暴(bao)龍(long)(long)(long)(long)(long)和霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)顴(quan)骨升突外側(ce)(ce)面都具(ju)有(you)一(yi)不(bu)(bu)甚(shen)明(ming)顯(xian)的(de)(de)(de)凹陷(xian),但(dan)是(shi)霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)這一(yi)凹陷(xian)明(ming)顯(xian)小(xiao)于(yu)特暴(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)。兩者的(de)(de)(de)顴(quan)骨后突都具(ju)有(you)二分支(zhi)的(de)(de)(de)結構,區別在于(yu)特暴(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)顴(quan)骨后突腹(fu)(fu)側(ce)(ce)支(zhi)的(de)(de)(de)厚度不(bu)(bu)均勻,其腹(fu)(fu)緣較厚,向背緣發展(zhan)的(de)(de)(de)過程中逐漸變薄,而(er)霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)顴(quan)骨后突腹(fu)(fu)側(ce)(ce)支(zhi)的(de)(de)(de)厚度更(geng)為(wei)(wei)均一(yi),且外側(ce)(ce)面更(geng)為(wei)(wei)明(ming)顯(xian)的(de)(de)(de)隆起。特暴(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)方(fang)顴(quan)骨柄明(ming)顯(xian)比(bi)霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)更(geng)加纖弱。霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)方(fang)顴(quan)骨前突比(bi)特暴(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)更(geng)為(wei)(wei)寬大(da),能夠覆蓋住顴(quan)骨后突的(de)(de)(de)大(da)部(bu)分面積,且內(nei)側(ce)(ce)面具(ju)有(you)一(yi)個(ge)很大(da)的(de)(de)(de)凹陷(xian)。特暴(bao)龍(long)(long)(long)(long)(long)顎骨上(shang)(shang)的(de)(de)(de)內(nei)鼻孔(kong)(kong)邊緣發育的(de)(de)(de)很明(ming)顯(xian),而(er)這一(yi)特征在霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)顎骨上(shang)(shang)相(xiang)(xiang)對(dui)較弱。特暴(bao)龍(long)(long)(long)(long)(long)外翼骨上(shang)(shang)的(de)(de)(de)氣(qi)孔(kong)(kong)比(bi)霸(ba)王(wang)(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)更(geng)小(xiao)。
特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)犁骨(gu)(gu)前突(tu)向前極度(du)延(yan)伸,與前頜(he)骨(gu)(gu)的(de)(de)(de)(de)(de)顎突(tu)相(xiang)接,而這(zhe)一特(te)征在(zai)霸(ba)王(wang)(wang)龍(long)(long)中不明顯。特(te)暴(bao)龍(long)(long)和霸(ba)王(wang)(wang)龍(long)(long)翼(yi)骨(gu)(gu)的(de)(de)(de)(de)(de)最大(da)區別在(zai)于顎骨(gu)(gu)關節面的(de)(de)(de)(de)(de)前邊緣。霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)翼(yi)骨(gu)(gu)額骨(gu)(gu)突(tu)的(de)(de)(de)(de)(de)前邊緣是一條斜線,而特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)翼(yi)骨(gu)(gu)顎骨(gu)(gu)突(tu)前邊緣向前突(tu)出(chu)。特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)上枕(zhen)骨(gu)(gu)的(de)(de)(de)(de)(de)內外向寬度(du)小(xiao)于枕(zhen)髁(ke),而霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)上枕(zhen)骨(gu)(gu)更(geng)寬。從側面看,特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)齒(chi)(chi)骨(gu)(gu)吻(wen)端比霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)更(geng)加(jia)尖細一點。霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)夾板骨(gu)(gu)腹(fu)緣相(xiang)較于特(te)暴(bao)龍(long)(long)有更(geng)大(da)的(de)(de)(de)(de)(de)傾斜角(jiao)度(du),并且夾板骨(gu)(gu)的(de)(de)(de)(de)(de)前端位于更(geng)高的(de)(de)(de)(de)(de)位置。特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)上隅骨(gu)(gu)孔(kong)較霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)要發達。霸(ba)王(wang)(wang)龍(long)(long)和特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)成(cheng)體(ti)上頜(he)骨(gu)(gu)齒(chi)(chi)和齒(chi)(chi)骨(gu)(gu)齒(chi)(chi)的(de)(de)(de)(de)(de)數量不一樣。
歸(gui)入(ru)標本:PIN 551-2是(shi)一具接近(jin)完(wan)整的骨架。最早被歸(gui)入(ru)到特暴(bao)龍(long)屬的歸(gui)入(ru)種(zhong)埃夫(fu)雷莫夫(fu)特暴(bao)龍(long)(Tarbosaurus efremovi)。現已歸(gui)入(ru)到勇(yong)士(shi)特暴(bao)龍(long),且勇(yong)士(shi)特暴(bao)龍(long)是(shi)特暴(bao)龍(long)屬下的唯一有效(xiao)種(zhong)。
歸入標本(ben):PIN 553-1保存了頭骨,背(bei)椎和尾(wei)椎,掌骨和跖(zhi)骨。最(zui)開始被歸入到蛇發女怪(guai)龍(long)屬。現已歸入勇士特(te)暴(bao)龍(long)。
歸(gui)入(ru)標本:PIN 552-2保存了(le)頭骨(gu)和頭后骨(gu)骼。最開始被歸(gui)入(ru)到蛇發女(nv)怪龍(long)屬(shu)。現(xian)已歸(gui)入(ru)勇(yong)士特暴(bao)龍(long)。
歸入標本(ben):MPC-D 107/7是一件幼年的(de)標本(ben),關聯保存。除了(le)缺少頸椎(zhui),前部背椎(zhui)和末(mo)端尾(wei)椎(zhui),其它骨頭基本(ben)都(dou)有(you)保存。
歸入標本:ZPAL MgD-I/3幾乎完(wan)整(zheng)的(de)中等體型的(de)個體。保(bao)存了頭骨(gu),頸椎和(he)背(bei)椎,10節(jie)近端(duan)尾椎,完(wan)整(zheng)的(de)腸骨(gu),恥骨(gu)和(he)坐骨(gu),肋骨(gu)和(he)腹膜(mo)肋,完(wan)整(zheng)的(de)肩胛骨(gu),烏喙骨(gu),左側的(de)前肢,不完(wan)整(zheng)保(bao)存的(de)右側后肢。
歸(gui)入標本(ben):ZPAL Mgd-I/4部分保(bao)存(cun)的大(da)型個體。保(bao)存(cun)了左側后肢,腸骨,13節椎體。
歸入標本:ZPAL Mgd-I/5不(bu)完整保(bao)(bao)存的大型(xing)骨架(jia)。保(bao)(bao)存了左(zuo)側(ce)上頜骨,左(zuo)側(ce)方骨,左(zuo)側(ce)的下(xia)頜和(he)右側(ce)的下(xia)頜碎片(pian),11節(jie)左(zuo)側(ce)肋(lei)骨的碎片(pian),恥(chi)骨碎片(pian),坐骨,腸骨碎片(pian),左(zuo)側(ce)后肢和(he)右側(ce)跖骨,和(he)很多碎屑。
歸入標本:ZPAL Mgd-I/26不完整保存的上(shang)頜(he)骨(gu)和一部分保存下(xia)來的牙齒。
歸入標本:ZPAL Mgd-I/29不完整(zheng)(zheng)的大型骨(gu)架。保存(cun)了頭(tou)骨(gu)和(he)較為(wei)完整(zheng)(zheng)的齒列。頭(tou)后骨(gu)骼保存(cun)了6節頸椎(zhui),5節薦椎(zhui),22節尾(wei)椎(zhui),11節右側(ce)肋骨(gu),腸骨(gu),不完整(zheng)(zheng)的恥骨(gu)和(he)近端(duan)坐骨(gu),左側(ce)肱骨(gu),尺骨(gu)和(he)橈骨(gu)的遠端(duan),第一手指,幾乎完整(zheng)(zheng)的右側(ce)后肢。
歸入標本:ZPAL Mgd-I/31右(you)側上(shang)頜骨的近端。
歸(gui)入標本(ben):ZPAL Mgd-I/34頭骨(gu)右側的(de)碎(sui)片。
歸入標本:ZPAL Mgd-I/38不完全(quan)保存的(de)頭骨(gu),12節(jie)(jie)肋(lei)骨(gu),右側(ce)(ce)股骨(gu)的(de)遠(yuan)端(duan)。右側(ce)(ce)脛骨(gu)的(de)遠(yuan)端(duan),右側(ce)(ce)的(de)3-4跖骨(gu),第4趾的(de)近端(duan)趾節(jie)(jie)骨(gu)。
歸入標本:ZPAL Mgd-I/44頭骨的右側。
歸入標(biao)本:ZPAL Mgd-I/45頭骨碎片(pian)。
歸入標(biao)本(ben):ZPAL Mgd-I/46右側(ce)下頜碎片和(he)(he)其它7塊(kuai)頭骨碎片,和(he)(he)兩節斷掉的背椎(zhui)肋(lei)骨。
歸入標本(ben):ZPAL Mgd-I/52左側下頜齒(chi)骨。
歸入標本:ZPAL Mgd-I/67右側顴骨。
歸入(ru)標本(ben):ZPAL Mgd-I/93頭骨(gu)腦顱(lu)。
歸入標(biao)本:ZPAL Mgd-I/109大(da)的(de)帶(dai)有(you)圍巖的(de)頭(tou)骨。
歸(gui)入標本:ZPAL Mgd-I/178頭骨(gu)的碎片,脊椎骨(gu)和股骨(gu)。
歸入(ru)標本:GIN 100/61頭骨碎片和頭后骨骼。
歸入(ru)標本:GIN 100/62頭(tou)骨碎片和頭(tou)后骨骼。
歸(gui)入標(biao)本:GIN 100/65頭骨(gu)的右(you)半部分。
歸入標本:GIN 100/67頭(tou)骨的碎片,腦顱。
歸入標本:GIN 100/69枕區。
歸入(ru)標本:GIN 100/70頭骨碎(sui)片和(he)椎體。
歸(gui)入標本:GIN 107/2完整的骨架。
歸(gui)入標本:GIN 107/3頭骨。
歸(gui)(gui)入(ru)標(biao)本:PIN 551-3骨架,保存狀況未描述。最早(zao)被歸(gui)(gui)入(ru)到特(te)暴(bao)龍屬(shu)的(de)歸(gui)(gui)入(ru)種埃夫雷莫夫特(te)暴(bao)龍。現已歸(gui)(gui)入(ru)到勇士特(te)暴(bao)龍。
歸(gui)入(ru)標本(ben):PIN 551-4骨架,保存狀況未(wei)描述(shu)。最早被歸(gui)入(ru)到特(te)(te)暴(bao)龍(long)屬的歸(gui)入(ru)種埃夫雷莫(mo)夫特(te)(te)暴(bao)龍(long)。現已歸(gui)入(ru)到勇士特(te)(te)暴(bao)龍(long)。
歸入標(biao)本:PIN 551-91右側上頜(he)骨。最早被歸入到(dao)特(te)(te)暴龍屬的歸入種埃夫雷莫夫特(te)(te)暴龍。現已歸入到(dao)勇士特(te)(te)暴龍。
歸(gui)入(ru)標本:PIN 552-1目前僅有(you)復制品保(bao)存。最(zui)早被歸(gui)入(ru)到(dao)特(te)暴龍屬的歸(gui)入(ru)種(zhong)埃夫雷莫(mo)夫特(te)暴龍。現已(yi)歸(gui)入(ru)到(dao)勇士特(te)暴龍。
歸(gui)(gui)(gui)(gui)入(ru)標本:PIN 553-2保存狀況未描(miao)述。早被歸(gui)(gui)(gui)(gui)入(ru)到特暴龍屬的(de)歸(gui)(gui)(gui)(gui)入(ru)種埃(ai)夫雷莫夫特暴龍。現已歸(gui)(gui)(gui)(gui)入(ru)到勇士特暴龍。
歸(gui)入標本:IVPP V 4878火焰山鄯善龍(long)(long)(Shanshanosaurus huoyanshanensis)的正(zheng)型標本,發現(xian)于中國新疆(jiang),現(xian)在認為(wei)可能(neng)是(shi)特暴龍(long)(long)的幼體。保(bao)存了上頜骨(gu),下(xia)頜大部分骨(gu)塊,多節頸椎,背椎和薦(jian)椎,肩帶骨(gu)骼(ge),肱骨(gu),恥骨(gu)遠端,股骨(gu)和脛骨(gu)近端。
歸入標(biao)本(ben):IVPP V 4733欒川暴龍(Tyrannosaurus luanchuanensis)的正型標(biao)本(ben),為(wei)5顆(ke)牙齒(chi),現(xian)認為(wei)也可能屬(shu)于特暴龍。
歸入標(biao)本:IVPP V 836為破(po)碎(sui)金剛口龍(Chingkankousaurus fragilis)的正(zheng)型標(biao)本,為一節不(bu)完整(zheng)的肩(jian)胛骨標(biao)本。布魯(lu)薩特等人認為很有可能(neng)是特暴龍,但不(bu)太確定。
特暴龍(long)(long)(long)同(tong)霸王(wang)龍(long)(long)(long)一(yi)樣,是暴龍(long)(long)(long)超(chao)科中最進步(bu)的(de)成(cheng)員(yuan)。大多數的(de)系(xi)統(tong)發育分析都支持霸王(wang)龍(long)(long)(long)和特暴龍(long)(long)(long)構(gou)成(cheng)一(yi)個姐妹(mei)群(qun)(qun),諸(zhu)城暴龍(long)(long)(long)作為(wei)(wei)兩者(zhe)所構(gou)成(cheng)單(dan)系(xi)的(de)姐妹(mei)群(qun)(qun)。也有(you)一(yi)些分析認為(wei)(wei)特暴龍(long)(long)(long)與諸(zhu)城暴龍(long)(long)(long)的(de)親(qin)緣關(guan)系(xi)更(geng)近。
如果不考慮(lv)標(biao)本(ben)(ben)量很(hen)少(shao)且質量很(hen)差(cha)的(de)(de)諸城暴(bao)(bao)龍(long),特暴(bao)(bao)龍(long)的(de)(de)體型基(ji)(ji)本(ben)(ben)穩坐(zuo)暴(bao)(bao)龍(long)類的(de)(de)第(di)二把(ba)交椅。最大(da)的(de)(de)特暴(bao)(bao)龍(long)頭骨(gu)長(chang)(chang)度達到1.3米(mi),這一大(da)小顯(xian)著的(de)(de)大(da)于其它的(de)(de)大(da)型暴(bao)(bao)龍(long)類,并基(ji)(ji)本(ben)(ben)達到了個別霸(ba)王龍(long)成年(nian)標(biao)本(ben)(ben)的(de)(de)頭骨(gu)長(chang)(chang)度。目前最大(da)的(de)(de)特暴(bao)(bao)龍(long)標(biao)本(ben)(ben)體長(chang)(chang)是肯(ken)定能超過10米(mi)的(de)(de)。特暴(bao)(bao)龍(long)的(de)(de)體重一般估計(ji)在(zai)4噸左右。
通過(guo)(guo)仔細(xi)對比霸王龍(long)(long)(long)(long)及其它(ta)北(bei)(bei)(bei)美暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)亞(ya)科成員的(de)(de)(de)(de)(de)頭(tou)骨(gu)(gu)各個骨(gu)(gu)塊(kuai)骨(gu)(gu)縫與(yu)(yu)特(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)差異(yi)可以發(fa)現霸王龍(long)(long)(long)(long)與(yu)(yu)特(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)頭(tou)骨(gu)(gu)具(ju)有(you)不太一(yi)(yi)樣的(de)(de)(de)(de)(de)生物力(li)學結構(gou)(gou)。大型暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)類(lei)都(dou)具(ju)有(you)毀滅性的(de)(de)(de)(de)(de)咬合(he)力(li),而(er)它(ta)們(men)的(de)(de)(de)(de)(de)頭(tou)骨(gu)(gu)也(ye)必須在撕咬獵物的(de)(de)(de)(de)(de)同(tong)時承受同(tong)樣巨(ju)大的(de)(de)(de)(de)(de)反作用力(li)。北(bei)(bei)(bei)美洲的(de)(de)(de)(de)(de)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)亞(ya)科主要通過(guo)(guo)上頜骨(gu)(gu)-鼻骨(gu)(gu)-淚(lei)(lei)骨(gu)(gu)-額骨(gu)(gu)/前額骨(gu)(gu)的(de)(de)(de)(de)(de)緊(jin)(jin)密關(guan)(guan)節(jie)構(gou)(gou)成來承受撕咬獵物過(guo)(guo)程中(zhong)(zhong)產生的(de)(de)(de)(de)(de)反作用力(li)。其中(zhong)(zhong),鼻骨(gu)(gu)與(yu)(yu)淚(lei)(lei)骨(gu)(gu)構(gou)(gou)成了非常強的(de)(de)(de)(de)(de)關(guan)(guan)節(jie),且這(zhe)一(yi)(yi)特(te)征普遍(bian)存在于北(bei)(bei)(bei)美的(de)(de)(de)(de)(de)大型暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)亞(ya)科中(zhong)(zhong)。對于特(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)研究顯示,與(yu)(yu)北(bei)(bei)(bei)美洲的(de)(de)(de)(de)(de)親戚們(men)相(xiang)比,特(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)淚(lei)(lei)骨(gu)(gu)前突向下偏轉的(de)(de)(de)(de)(de)更多,并且擁(yong)有(you)比北(bei)(bei)(bei)美洲暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)類(lei)更為強壯(zhuang)的(de)(de)(de)(de)(de)上頜骨(gu)(gu)升突,使得(de)這(zhe)兩個塊(kuai)骨(gu)(gu)頭(tou)通過(guo)(guo)復雜的(de)(de)(de)(de)(de)關(guan)(guan)節(jie)面緊(jin)(jin)密關(guan)(guan)節(jie)在一(yi)(yi)起。同(tong)時特(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)前額骨(gu)(gu)與(yu)(yu)淚(lei)(lei)骨(gu)(gu)也(ye)有(you)非常緊(jin)(jin)密的(de)(de)(de)(de)(de)關(guan)(guan)節(jie)。非常強大的(de)(de)(de)(de)(de)上頜骨(gu)(gu)-淚(lei)(lei)骨(gu)(gu)關(guan)(guan)節(jie)也(ye)見于異(yi)特(te)龍(long)(long)(long)(long)和(he)中(zhong)(zhong)華盜龍(long)(long)(long)(long)這(zhe)樣的(de)(de)(de)(de)(de)大型異(yi)特(te)龍(long)(long)(long)(long)類(lei)中(zhong)(zhong)。
對(dui)于(yu)異特(te)龍(long)(long)(long)(long)頭(tou)骨(gu)的(de)(de)(de)(de)(de)有限元分(fen)析顯示,異特(te)龍(long)(long)(long)(long)在攻擊獵物時(shi),頭(tou)骨(gu)的(de)(de)(de)(de)(de)上頜(he)(he)(he)骨(gu)-淚骨(gu)關(guan)節(jie)處會(hui)承受(shou)非常大(da)(da)的(de)(de)(de)(de)(de)反(fan)作用力。另外(wai)一(yi)點區別是,北美洲(zhou)的(de)(de)(de)(de)(de)暴(bao)龍(long)(long)(long)(long)類(lei)(lei)下(xia)(xia)頜(he)(he)(he)的(de)(de)(de)(de)(de)齒(chi)骨(gu)和(he)后(hou)面的(de)(de)(de)(de)(de)骨(gu)塊具有較為靈活的(de)(de)(de)(de)(de)關(guan)節(jie),使得它(ta)們(men)的(de)(de)(de)(de)(de)下(xia)(xia)頜(he)(he)(he)可以一(yi)定程度的(de)(de)(de)(de)(de)內外(wai)擴(kuo)展(zhan)和(he)收縮,而特(te)暴(bao)龍(long)(long)(long)(long)沒有這樣(yang)的(de)(de)(de)(de)(de)關(guan)節(jie),使得它(ta)們(men)的(de)(de)(de)(de)(de)下(xia)(xia)頜(he)(he)(he)更為一(yi)體(ti)化(hua)。造成這些區別的(de)(de)(de)(de)(de)原因主(zhu)要(yao)(yao)是亞(ya)(ya)洲(zhou)暴(bao)龍(long)(long)(long)(long)類(lei)(lei)和(he)北美洲(zhou)的(de)(de)(de)(de)(de)暴(bao)龍(long)(long)(long)(long)類(lei)(lei)所面對(dui)的(de)(de)(de)(de)(de)獵物不(bu)太一(yi)樣(yang)。晚白堊世北美洲(zhou)的(de)(de)(de)(de)(de)暴(bao)龍(long)(long)(long)(long)類(lei)(lei)主(zhu)要(yao)(yao)捕食角(jiao)龍(long)(long)(long)(long)類(lei)(lei)和(he)鴨嘴(zui)龍(long)(long)(long)(long)類(lei)(lei),而亞(ya)(ya)洲(zhou)的(de)(de)(de)(de)(de)暴(bao)龍(long)(long)(long)(long)類(lei)(lei)則(ze)有更多(duo)的(de)(de)(de)(de)(de)機會(hui)捕獵大(da)(da)型(xing)蜥腳(jiao)類(lei)(lei)恐龍(long)(long)(long)(long),比(bi)如泰(tai)坦巨龍(long)(long)(long)(long)類(lei)(lei)。目(mu)前還未有研究(jiu)使用模(mo)型(xing)化(hua)的(de)(de)(de)(de)(de)方法(fa)精確計(ji)算過特(te)暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)咬合力,但定性(xing)的(de)(de)(de)(de)(de)比(bi)較認為,特(te)暴(bao)龍(long)(long)(long)(long)與其它(ta)大(da)(da)型(xing)暴(bao)龍(long)(long)(long)(long)類(lei)(lei)一(yi)樣(yang)擁(yong)有非常巨大(da)(da)的(de)(de)(de)(de)(de)咬合力。
菲利普(pu).柯瑞等人在2003年的(de)一項研(yan)究中報(bao)道了(le)2個可能屬于(yu)特暴龍(long)的(de)腳(jiao)印(yin)。這些腳(jiao)印(yin)非常巨(ju)大(da),有61厘米(mi)長,保(bao)存了(le)第(di)三趾(zhi)爪在地上留(liu)下的(de)痕(hen)(hen)跡(ji)。同時放大(da)觀察腳(jiao)印(yin)還(huan)可以(yi)看到里面的(de)皮(pi)膚(fu)印(yin)痕(hen)(hen)。皮(pi)膚(fu)印(yin)痕(hen)(hen)中的(de)每個鱗(lin)片大(da)約(yue)有2毫米(mi)寬(kuan)。另(ling)外一件特暴龍(long)的(de)骨架上也發現了(le)皮(pi)膚(fu)印(yin)痕(hen)(hen)。皮(pi)膚(fu)印(yin)痕(hen)(hen)保(bao)留(liu)在接(jie)近咽喉的(de)位置,鱗(lin)片的(de)寬(kuan)度約(yue)為2.4毫米(mi)。遺憾(han)的(de)是這件標本已經被破壞,無法對(dui)其進行(xing)更深入的(de)研(yan)究。
一(yi)件(jian)特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)標本保存了完整的(de)(de)(de)顱腔(qiang)。通(tong)過制作顱內模(mo)可以了解(jie)特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)腦(nao)(nao)(nao)結(jie)構(gou)。特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)腦(nao)(nao)(nao)結(jie)構(gou)與霸王龍(long)(long)(long)十分相似,其主要(yao)的(de)(de)(de)區別(bie)僅(jin)存在于(yu)個別(bie)腦(nao)(nao)(nao)神(shen)經基部(bu)的(de)(de)(de)位置,如三叉神(shen)經和副神(shen)經。一(yi)只12米(mi)長的(de)(de)(de)特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)腦(nao)(nao)(nao)體積約為(wei)184立方厘米(mi)。特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)和霸王龍(long)(long)(long)一(yi)樣擁有非常大的(de)(de)(de)嗅(xiu)葉(xie),這(zhe)說明它(ta)們有良好的(de)(de)(de)嗅(xiu)覺。同時它(ta)們也(ye)具有發達的(de)(de)(de)犁鼻器用(yong)于(yu)探測荷爾蒙,這(zhe)說明特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)可能擁有復雜的(de)(de)(de)交配(pei)行為(wei)。特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)聽(ting)神(shen)經也(ye)很發達,說明它(ta)們有較好的(de)(de)(de)聽(ting)力。特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)中(zhong)腦(nao)(nao)(nao)頂部(bu)(midbrain tectum)不甚發達,它(ta)的(de)(de)(de)動眼神(shen)經和視神(shen)經也(ye)是如此。在頭骨的(de)(de)(de)宏觀(guan)形態形態上,特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)頭骨后端(duan)向外(wai)側(ce)擴展(zhan)的(de)(de)(de)較少(shao),其雙眼視覺疊加范圍不如霸王龍(long)(long)(long)。因此特(te)(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)在生(sheng)活(huo)的(de)(de)(de)時候對嗅(xiu)覺和聽(ting)覺的(de)(de)(de)依(yi)賴(lai)(lai)要(yao)遠大于(yu)對視覺的(de)(de)(de)依(yi)賴(lai)(lai)。
同(tong)北美的(de)(de)(de)(de)(de)(de)大(da)型暴(bao)(bao)龍(long)(long)類一樣,特(te)暴(bao)(bao)龍(long)(long)在生長發(fa)育過程(cheng)中(zhong)也伴隨著(zhu)一系列的(de)(de)(de)(de)(de)(de)形態變(bian)(bian)化(hua)。目(mu)前發(fa)現(xian)的(de)(de)(de)(de)(de)(de)大(da)多數(shu)(shu)特(te)暴(bao)(bao)龍(long)(long)標本屬于(yu)(yu)亞成(cheng)(cheng)年(nian)個(ge)(ge)體(ti)或成(cheng)(cheng)年(nian)個(ge)(ge)體(ti),只有(you)很少(shao)的(de)(de)(de)(de)(de)(de)幼(you)(you)(you)年(nian)個(ge)(ge)體(ti)。2011年(nian)發(fa)表的(de)(de)(de)(de)(de)(de)一件特(te)暴(bao)(bao)龍(long)(long)幼(you)(you)(you)體(ti)標本使得古生物(wu)學家(jia)對于(yu)(yu)特(te)暴(bao)(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)幼(you)(you)(you)年(nian)時期有(you)了更(geng)多了解。這(zhe)只幼(you)(you)(you)體(ti)在死亡的(de)(de)(de)(de)(de)(de)時候(hou)大(da)約只有(you)2-3歲。與成(cheng)(cheng)年(nian)個(ge)(ge)體(ti)相比,幼(you)(you)(you)體(ti)的(de)(de)(de)(de)(de)(de)頭骨(gu)較為(wei)脆弱,牙齒(chi)也不(bu)夠強(qiang)壯,說明幼(you)(you)(you)體(ti)的(de)(de)(de)(de)(de)(de)獵物(wu)與成(cheng)(cheng)體(ti)有(you)較大(da)的(de)(de)(de)(de)(de)(de)不(bu)同(tong)。與霸王龍(long)(long)和懼龍(long)(long)不(bu)同(tong)的(de)(de)(de)(de)(de)(de)是(shi),這(zhe)件幼(you)(you)(you)年(nian)特(te)暴(bao)(bao)龍(long)(long)標本的(de)(de)(de)(de)(de)(de)齒(chi)骨(gu)齒(chi)和上頜骨(gu)齒(chi)的(de)(de)(de)(de)(de)(de)數(shu)(shu)量(liang)與成(cheng)(cheng)年(nian)個(ge)(ge)體(ti)的(de)(de)(de)(de)(de)(de)數(shu)(shu)量(liang)一致,說明特(te)暴(bao)(bao)龍(long)(long)在個(ge)(ge)體(ti)發(fa)育的(de)(de)(de)(de)(de)(de)過程(cheng)中(zhong)不(bu)存(cun)在牙齒(chi)數(shu)(shu)量(liang)變(bian)(bian)化(hua)的(de)(de)(de)(de)(de)(de)現(xian)象。這(zhe)一區別是(shi)否廣泛存(cun)在與亞洲(zhou)和北美洲(zhou)的(de)(de)(de)(de)(de)(de)其它(ta)暴(bao)(bao)龍(long)(long)亞科物(wu)種還(huan)需要(yao)進一步檢驗。對于(yu)(yu)幼(you)(you)(you)體(ti)鞏膜環的(de)(de)(de)(de)(de)(de)研究表明,幼(you)(you)(you)體(ti)可能具有(you)夜(ye)行(xing)的(de)(de)(de)(de)(de)(de)習(xi)性。成(cheng)(cheng)年(nian)特(te)暴(bao)(bao)龍(long)(long)是(shi)否具有(you)夜(ye)行(xing)習(xi)性還(huan)需要(yao)更(geng)多的(de)(de)(de)(de)(de)(de)化(hua)石證據支(zhi)持。
特暴龍(long)(long)(long)(long)是其(qi)所處生態系(xi)統中最大(da)的捕食(shi)者,與其(qi)共存的植(zhi)食(shi)性恐龍(long)(long)(long)(long)包(bao)括蜥(xi)腳類(lei)(lei)的耐梅蓋(gai)特龍(long)(long)(long)(long)和后凹尾龍(long)(long)(long)(long),甲龍(long)(long)(long)(long)類(lei)(lei)的美甲龍(long)(long)(long)(long),腫頭龍(long)(long)(long)(long)類(lei)(lei)的傾頭龍(long)(long)(long)(long),鴨嘴(zui)龍(long)(long)(long)(long)類(lei)(lei)的櫛龍(long)(long)(long)(long)和巴(ba)思(si)缽氏龍(long)(long)(long)(long),屬(shu)(shu)于鐮(lian)刀(dao)龍(long)(long)(long)(long)類(lei)(lei)的鐮(lian)刀(dao)龍(long)(long)(long)(long),屬(shu)(shu)于似(si)鳥(niao)龍(long)(long)(long)(long)類(lei)(lei)似(si)鵝龍(long)(long)(long)(long),似(si)雞龍(long)(long)(long)(long)和體型巨大(da)的恐手龍(long)(long)(long)(long)。肉食(shi)性恐龍(long)(long)(long)(long)則包(bao)括同(tong)屬(shu)(shu)于暴龍(long)(long)(long)(long)亞科的分支龍(long)(long)(long)(long),屬(shu)(shu)于傷齒龍(long)(long)(long)(long)類(lei)(lei)的無聊(liao)龍(long)(long)(long)(long),鴕鳥(niao)龍(long)(long)(long)(long)和扎(zha)納巴(ba)扎(zha)爾(er)龍(long)(long)(long)(long),屬(shu)(shu)于竊(qie)蛋龍(long)(long)(long)(long)類(lei)(lei)的單足(zu)龍(long)(long)(long)(long),耐梅蓋(gai)特母龍(long)(long)(long)(long)和瑞欽龍(long)(long)(long)(long)。
特暴龍(long)曾登(deng)場于BBC的紀錄(lu)片(pian)《恐龍(long)兇(xiong)面目》以及(ji)《與龍(long)同行特輯:尋爪記(ji)》。近(jin)期它又在紀錄(lu)片(pian)《史前星球》中(zhong)登(deng)場。
在電影(ying)方面(mian),特暴龍(long)登場(chang)于(yu)韓國的動畫電影(ying)《韓半島的恐龍(long)》以及國產恐龍(long)動畫電影(ying)《恐龍(long)王》。
特暴龍位于食物鏈的頂端,是(shi)(shi)(shi)一種(zhong)頂(ding)級掠食動(dong)物。特(te)(te)暴(bao)(bao)龍(long)(long)(long)在暴(bao)(bao)龍(long)(long)(long)科中(zhong)的(de)分類位(wei)置仍未確定。以(yi)前,有些(xie)科學家認為(wei)勇士(shi)特(te)(te)暴(bao)(bao)龍(long)(long)(long)其實是(shi)(shi)(shi)北美洲(zhou)(zhou)暴(bao)(bao)龍(long)(long)(long)的(de)亞(ya)洲(zhou)(zhou)種(zhong)。如果屬(shu)實,將(jiang)使特(te)(te)暴(bao)(bao)龍(long)(long)(long)成為(wei)無(wu)效(xiao)的(de)分類。即使特(te)(te)暴(bao)(bao)龍(long)(long)(long)與暴(bao)(bao)龍(long)(long)(long)不(bu)是(shi)(shi)(shi)同種(zhong)動(dong)物,它們(men)被(bei)認為(wei)有接近的(de)親緣(yuan)關系。有些(xie)科學家認為(wei),同樣發現(xian)于蒙(meng)古(gu)的(de)分支龍(long)(long)(long),是(shi)(shi)(shi)特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)近親。
特暴龍(long)的化石記錄保存良好,已有數十個標本,包含至少5個完整的頭顱骨與骨骸。這(zhe)些化石讓科學家得以研究(jiu)它(ta)們的種系發生學、頭部力學、以及腦部結(jie)構。
特暴龍(long)(long)屬于暴龍(long)(long)科(ke)的(de)(de)暴龍(long)(long)亞(ya)(ya)(ya)科(ke)。該亞(ya)(ya)(ya)科(ke)還包含較(jiao)(jiao)早(zao)期的(de)(de)懼龍(long)(long)、較(jiao)(jiao)晚期的(de)(de)暴龍(long)(long),都發現于北美洲,可能(neng)還有(you)蒙古的(de)(de)分支龍(long)(long)。暴龍(long)(long)亞(ya)(ya)(ya)科(ke)包含親緣關系較(jiao)(jiao)接近暴龍(long)(long),而離艾伯(bo)塔(ta)龍(long)(long)較(jiao)(jiao)遠的(de)(de)物(wu)種;與(yu)艾伯(bo)塔(ta)龍(long)(long)亞(ya)(ya)(ya)科(ke)相(xiang)比,暴龍(long)(long)亞(ya)(ya)(ya)科(ke)的(de)(de)體格(ge)較(jiao)(jiao)重型,頭顱骨的(de)(de)比例較(jiao)(jiao)大,以及較(jiao)(jiao)長的(de)(de)股骨。
勇士特暴龍最初被視為暴龍的一個種,某些近年的分類也支持這個說法(fa)。其(qi)(qi)他的(de)科學家則(ze)將它(ta)們(men)列為獨(du)立的(de)屬,并(bing)為暴(bao)龍(long)(long)(long)(long)的(de)姐妹分(fen)類(lei)單(dan)元。在(zai)2003年,一個親緣分(fen)支(zhi)分(fen)類(lei)法(fa)研究提出分(fen)支(zhi)龍(long)(long)(long)(long)是特暴(bao)龍(long)(long)(long)(long)的(de)近(jin)親,因為它(ta)們(men)具有(you)其(qi)(qi)他暴(bao)龍(long)(long)(long)(long)亞科沒(mei)有(you)的(de)頭部特征。如果屬實,將排除(chu)特暴(bao)龍(long)(long)(long)(long)是暴(bao)龍(long)(long)(long)(long)的(de)一個異名(ming)的(de)可(ke)能性(xing),并(bing)顯示暴(bao)龍(long)(long)(long)(long)亞科在(zai)北美洲(zhou)與(yu)亞洲(zhou)演化出個別的(de)支(zhi)系。分(fen)支(zhi)龍(long)(long)(long)(long)的(de)唯一標本具有(you)幼(you)年體的(de)特征,但牙齒(chi)數量較(jiao)多,約76到78顆,而(er)且口鼻部上(shang)面有(you)獨(du)特的(de)低(di)矮骨質(zhi)瘤(liu),因此(ci)并(bing)非特暴(bao)龍(long)(long)(long)(long)的(de)幼(you)年體。
特(te)暴(bao)龍(long)過去生(sheng)存于潮濕的(de)泛濫(lan)平(ping)原,布滿者河道。在中(zhong)(zhong)國(guo)的(de)分布包(bao)括黑龍(long)江,河南(nan),山(shan)東,廣東,云南(nan),內蒙古等地(di),國(guo)外(wai)主要是蒙古。在中(zhong)(zhong)國(guo)黑龍(long)江,河南(nan),山(shan)東,廣東的(de)所謂中(zhong)(zhong)國(guo)"暴(bao)龍(long)"其實都是特(te)暴(bao)龍(long)。
特暴龍(long)(long)(long)是(shi)最大(da)(da)型(xing)(xing)(xing)的(de)(de)(de)暴龍(long)(long)(long)科動(dong)物(wu)之一,但略小于暴龍(long)(long)(long)。已知最大(da)(da)型(xing)(xing)(xing)的(de)(de)(de)個(ge)體(ti)(ti)身(shen)長12米(mi),頭部離(li)地(di)面約(yue)4.2米(mi)。一般體(ti)(ti)重3噸到5噸,最大(da)(da)的(de)(de)(de)化石體(ti)(ti)重可(ke)達到7.5噸。如同大(da)(da)部分已知的(de)(de)(de)暴龍(long)(long)(long)科恐龍(long)(long)(long),特暴龍(long)(long)(long)是(shi)種大(da)(da)型(xing)(xing)(xing)、二足(zu)掠(lve)食動(dong)物(wu),重達數噸,擁有(you)數十顆大(da)(da)型(xing)(xing)(xing)、銳利的(de)(de)(de)牙齒。特暴龍(long)(long)(long)的(de)(de)(de)下頜(he)有(you)特殊(shu)的(de)(de)(de)接合構造。另外(wai),就(jiu)前肢/身(shen)體(ti)(ti)比例而言(yan),特暴龍(long)(long)(long)擁有(you)暴龍(long)(long)(long)科中(zhong)最小型(xing)(xing)(xing)的(de)(de)(de)前肢。特暴龍(long)(long)(long)是(shi)種大(da)(da)型(xing)(xing)(xing)、二足(zu)掠(lve)食動(dong)物(wu),重達數噸。特暴龍(long)(long)(long)唯一一個(ge)有(you)效種的(de)(de)(de)體(ti)(ti)型(xing)(xing)(xing):勇士特暴龍(long)(long)(long)Tarbosaurus bataar 8-12米(mi)。
暴龍科(ke)的身體外形差異不大。特暴龍的頸部為(wei)S狀彎曲,其余的脊柱,包含尾巴(ba),與地(di)面保持(chi)者水平的姿態。
就前肢/身體(ti)比例而(er)言,特暴龍(long)(long)擁有(you)暴龍(long)(long)科中最小型(xing)的前肢。有(you)兩(liang)根迷你的手指。后肢長(chang)而(er)粗(cu)厚,將身體(ti)支撐(cheng)為二足(zu)的步態,上有(you)三(san)根腳趾。長(chang)而(er)重(zhong)(zhong)(zhong)的尾(wei)巴可(ke)以平(ping)衡頭部與(yu)胸(xiong)部的重(zhong)(zhong)(zhong)量,將重(zhong)(zhong)(zhong)心保持(chi)在臀部。
在2003年,特(te)(te)暴(bao)(bao)龍(long)(long)的(de)(de)顱骨(gu)(gu)(gu)首(shou)次經(jing)過完(wan)整(zheng)的(de)(de)研究(jiu)。科(ke)學家們發(fa)現特(te)(te)暴(bao)(bao)龍(long)(long)與北美(mei)洲(zhou)暴(bao)(bao)龍(long)(long)科(ke)之間(jian)有幾個顯著的(de)(de)差異(yi)。這些差異(yi)與顱骨(gu)(gu)(gu)在咬(yao)合(he)(he)時,力(li)量(liang)的(de)(de)傳遞(di)有關。當特(te)(te)暴(bao)(bao)龍(long)(long)的(de)(de)上(shang)頜咬(yao)住物(wu)體時,力(li)量(liang)從(cong)上(shang)頜骨(gu)(gu)(gu)傳遞(di)到(dao)上(shang)頜周遭(zao)的(de)(de)顱骨(gu)(gu)(gu)。而北美(mei)洲(zhou)暴(bao)(bao)龍(long)(long)科(ke)咬(yao)合(he)(he)時,力(li)量(liang)從(cong)上(shang)頜骨(gu)(gu)(gu)傳遞(di)到(dao)口鼻部上(shang)方的(de)(de)固(gu)定鼻骨(gu)(gu)(gu),然后再(zai)經(jing)由鼻骨(gu)(gu)(gu)與淚骨(gu)(gu)(gu)間(jian)的(de)(de)骨(gu)(gu)(gu)質連結,傳遞(di)到(dao)淚骨(gu)(gu)(gu)。
特(te)(te)暴龍的鼻骨(gu)(gu)(gu)(gu)與(yu)淚(lei)骨(gu)(gu)(gu)(gu)間(jian)缺乏骨(gu)(gu)(gu)(gu)質的連結。但上頜(he)骨(gu)(gu)(gu)(gu)的后方有(you)個大型突起,楔合(he)(he)入淚(lei)骨(gu)(gu)(gu)(gu)內;而北美(mei)洲暴龍的上頜(he)骨(gu)(gu)(gu)(gu)后突很小(xiao)。這個特(te)(te)征顯示,咬合(he)(he)的力量從特(te)(te)暴龍的上頜(he)骨(gu)(gu)(gu)(gu)直(zhi)接傳遞到淚(lei)骨(gu)(gu)(gu)(gu)。而淚(lei)骨(gu)(gu)(gu)(gu)與(yu)額骨(gu)(gu)(gu)(gu)、前額骨(gu)(gu)(gu)(gu)之間(jian)更為(wei)牢固。由于(yu)上頜(he)骨(gu)(gu)(gu)(gu)、淚(lei)骨(gu)(gu)(gu)(gu)、額骨(gu)(gu)(gu)(gu)、前額骨(gu)(gu)(gu)(gu)之間(jian)牢牢地固定(ding)者,使得上頜(he)非(fei)常堅固。
另(ling)一個主要的差(cha)別則是堅(jian)固的下(xia)(xia)頜(he)。許多(duo)獸(shou)腳類(lei)恐龍(long)(long),包括北美洲暴(bao)龍(long)(long)科在內(nei),下(xia)(xia)頜(he)的齒骨與后(hou)面骨頭間有靈活(huo)的關節。特(te)暴(bao)龍(long)(long)的隅骨側(ce)邊棱脊連接者齒骨后(hou)方的方形突(tu),使(shi)它們的下(xia)(xia)頜(he)無法靈活(huo)外內(nei)扳動。
有些科(ke)學(xue)(xue)家提出假設,認為(wei)特(te)(te)暴龍(long)的(de)堅硬頭部是(shi)種(zhong)適(shi)應演化,用來獵殺耐梅(mei)蓋(gai)特(te)(te)組(zu)的(de)大型(xing)蜥(xi)腳(jiao)類(lei)恐龍(long),泰坦巨(ju)龍(long)類(lei),因為(wei)晚白堊紀的(de)北(bei)美洲并沒有如此巨(ju)大的(de)恐龍(long)存在。這種(zhong)頭部力(li)學(xue)(xue)的(de)差異也影響了暴龍(long)科(ke)的(de)種(zhong)系發生學(xue)(xue)。同樣發現于蒙古的(de)分(fen)支龍(long),也具有類(lei)似(si)特(te)(te)暴龍(long)的(de)頭部力(li)學(xue)(xue)特(te)(te)征,這顯示(shi)暴龍(long)并非特(te)(te)暴龍(long)的(de)近親。特(te)(te)暴龍(long)與暴龍(long)的(de)相(xiang)似(si)處可(ke)能導(dao)因于它們巨(ju)大的(de)體型(xing),是(shi)平行演化的(de)結果。
一個(ge)在1948年發(fa)現的(de)顱(lu)骨,一度被歸類于(yu)蛇發(fa)女怪龍的(de)G. lancinator,因為具有顱(lu)腔(qiang),有助于(yu)科學家了(le)解特暴龍的(de)腦部(bu)結構。在1965年,馬(ma)列夫(fu)制(zhi)作了(le)一個(ge)石膏顱(lu)腔(qiang)模型(xing),并做了(le)初步的(de)腦部(bu)形狀檢(jian)查。在2005年,謝爾蓋·薩伐黎耶夫(fu)(Sergei V. Saveliev)制(zhi)作了(le)一個(ge)聚氨(an)酯(zhi)顱(lu)腔(qiang)模型(xing),并做了(le)更(geng)詳細的(de)腦部(bu)結構與功能研究。
根(gen)據暴(bao)龍(long)(long)(long)(long)(long)(long)的(de)(de)(de)腦(nao)部(bu)(bu)結構研究,它(ta)們(men)兩(liang)者的(de)(de)(de)腦(nao)部(bu)(bu)結構相似,只(zhi)有某些(xie)腦(nao)神(shen)(shen)(shen)經(jing)(jing)(jing)根(gen)部(bu)(bu)的(de)(de)(de)位置不一(yi)樣,包含三叉神(shen)(shen)(shen)經(jing)(jing)(jing)與(yu)副神(shen)(shen)(shen)經(jing)(jing)(jing)。暴(bao)龍(long)(long)(long)(long)(long)(long)科的(de)(de)(de)腦(nao)部(bu)(bu)較(jiao)類似鱷魚與(yu)其他爬蟲類,而較(jiao)不類似鳥類。特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)的(de)(de)(de)腦(nao)部(bu)(bu)體(ti)積為184立方厘(li)米(mi)。腦(nao)部(bu)(bu)的(de)(de)(de)大(da)型嗅球、末端神(shen)(shen)(shen)經(jing)(jing)(jing)、嗅神(shen)(shen)(shen)經(jing)(jing)(jing),顯(xian)示(shi)(shi)特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)具有靈(ling)敏的(de)(de)(de)嗅覺,這(zhe)點如同暴(bao)龍(long)(long)(long)(long)(long)(long)。犁鼻(bi)球大(da)型,而且與(yu)嗅球分離,顯(xian)示(shi)(shi)可(ke)以感應費洛蒙(meng)的(de)(de)(de)犁鼻(bi)器發展(zhan)良(liang)好(hao)(hao)。這(zhe)個特(te)(te)征顯(xian)示(shi)(shi)特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)可(ke)能具有復雜的(de)(de)(de)求(qiu)偶行為。聽神(shen)(shen)(shen)經(jing)(jing)(jing)也(ye)很(hen)大(da),顯(xian)示(shi)(shi)它(ta)們(men)的(de)(de)(de)聽力很(hen)好(hao)(hao),可(ke)能用在聲音的(de)(de)(de)溝通與(yu)警(jing)告上。聽神(shen)(shen)(shen)經(jing)(jing)(jing)連(lian)接者發展(zhan)良(liang)好(hao)(hao)的(de)(de)(de)前庭系統,顯(xian)示(shi)(shi)它(ta)們(men)的(de)(de)(de)平衡感與(yu)協調性(xing)很(hen)好(hao)(hao)。相反(fan)地,與(yu)視力有關的(de)(de)(de)腦(nao)部(bu)(bu)區域(yu)與(yu)神(shen)(shen)(shen)經(jing)(jing)(jing)較(jiao)小。爬蟲類的(de)(de)(de)中腦(nao)頂(ding)蓋(gai)連(lian)接控制眼(yan)球的(de)(de)(de)視神(shen)(shen)(shen)經(jing)(jing)(jing)與(yu)動眼(yan)神(shen)(shen)(shen)經(jing)(jing)(jing),是用來處理視力訊息,但特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)的(de)(de)(de)中腦(nao)頂(ding)蓋(gai)非常小。暴(bao)龍(long)(long)(long)(long)(long)(long)的(de)(de)(de)眼(yan)睛朝(chao)前,因此(ci)具有一(yi)定(ding)程度的(de)(de)(de)立體(ti)視覺;但特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)的(de)(de)(de)顱骨狹(xia)窄,眼(yan)睛朝(chao)向兩(liang)側,如同典型的(de)(de)(de)暴(bao)龍(long)(long)(long)(long)(long)(long)科動物。這(zhe)些(xie)特(te)(te)征顯(xian)示(shi)(shi)特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)較(jiao)依靠嗅覺與(yu)聽覺,而非視覺。
已(yi)知(zhi)最大(da)(da)(da)型的(de)特暴(bao)龍(long)(long)顱(lu)骨長度超過1.3米,小于暴(bao)龍(long)(long)和諸城暴(bao)龍(long)(long)以(yi)及懼龍(long)(long),但大(da)(da)(da)于其它的(de)暴(bao)龍(long)(long)科。如同暴(bao)龍(long)(long),特暴(bao)龍(long)(long)的(de)顱(lu)骨高大(da)(da)(da),前段(duan)狹(xia)窄。顱(lu)骨后段(duan)擴張幅度不大(da)(da)(da),意味著(zhu)特暴(bao)龍(long)(long)所以(yi)不如暴(bao)龍(long)(long)擁有(you)的(de)立體視覺好。
顱(lu)骨的大型洞孔可(ke)減輕重量。特(te)暴龍(long)的下頜有特(te)殊(shu)的接合構造。
顎部(bu)有60到(dao)64顆(ke)牙(ya)齒(chi)(chi),略(lve)少(shao)于(yu)暴(bao)(bao)龍(long)(long)(long),但(dan)大于(yu)其它(ta)(ta)體型(xing)較(jiao)小的(de)暴(bao)(bao)龍(long)(long)(long)科(ke)(ke),例如(ru)蛇發女(nv)怪龍(long)(long)(long)與分支龍(long)(long)(long)。大部(bu)分的(de)牙(ya)齒(chi)(chi)橫剖面為橢圓(yuan)形,而前上(shang)顎骨的(de)牙(ya)齒(chi)(chi)橫剖面為D形。暴(bao)(bao)龍(long)(long)(long)科(ke)(ke)都具有這種異型(xing)齒(chi)(chi)特(te)(te)征。上(shang)顎骨的(de)牙(ya)齒(chi)(chi)最(zui)長(chang)(chang),齒(chi)(chi)冠長(chang)(chang)達85公厘(li)。如(ru)同其它(ta)(ta)大型(xing)暴(bao)(bao)龍(long)(long)(long)科(ke)(ke)與現(xian)代的(de)科(ke)(ke)莫(mo)多(duo)龍(long)(long)(long),特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)幼年與近成(cheng)年個體的(de)身上(shang)發現(xian)過有齒(chi)(chi)痕,符合成(cheng)年個體或其它(ta)(ta)較(jiao)小型(xing)獸(shou)腳(jiao)類恐龍(long)(long)(long)的(de)牙(ya)齒(chi)(chi)。
特(te)暴龍(long)與分支龍(long)的(de)下(xia)顎外側各有一道棱脊,從隅骨(gu)延(yan)伸(shen)到齒骨(gu)后(hou)方,形成相扣的(de)結構。其它(ta)暴龍(long)科動(dong)物(wu)缺乏這道棱脊,因(yin)此(ci)下(xia)顎更為靈活(huo)。
特(te)暴龍(long)(long)位于食(shi)物鏈的頂(ding)端,是一(yi)種(zhong)頂(ding)級掠食(shi)者(zhe),可能以大型恐龍(long)(long)為(wei)食(shi),例如鴨嘴龍(long)(long)類的櫛(zhi)龍(long)(long),或是蜥腳類的納(na)摩蓋(gai)吐龍(long)(long)。
成年特(te)暴(bao)龍(long)(long)(long)(long)可能與其(qi)它(ta)(ta)小(xiao)(xiao)型獸腳(jiao)類恐(kong)龍(long)(long)(long)(long)有(you)少許競(jing)爭,例如傷齒龍(long)(long)(long)(long)科(ke)的無聊龍(long)(long)(long)(long)、鴕鳥龍(long)(long)(long)(long)、蜥鳥龍(long)(long)(long)(long),以(yi)(yi)及偷蛋龍(long)(long)(long)(long)下目(mu)(mu)的單足(zu)龍(long)(long)(long)(long)、瑞欽(qin)龍(long)(long)(long)(long),或者(zhe)還有(you)小(xiao)(xiao)掠龍(long)(long)(long)(long),一(yi)種有(you)時被(bei)認為是基(ji)底暴(bao)龍(long)(long)(long)(long)超科(ke)的恐(kong)龍(long)(long)(long)(long)。其(qi)它(ta)(ta)的獸腳(jiao)類恐(kong)龍(long)(long)(long)(long),包含(han):巨大的鐮刀龍(long)(long)(long)(long)、似鳥龍(long)(long)(long)(long)下目(mu)(mu)的似鵝龍(long)(long)(long)(long)、似雞龍(long)(long)(long)(long)、恐(kong)手龍(long)(long)(long)(long),鐮刀龍(long)(long)(long)(long)可能是草食(shi)性,而上述似鳥龍(long)(long)(long)(long)類恐(kong)龍(long)(long)(long)(long)可能為雜食(shi)性,以(yi)(yi)小(xiao)(xiao)型動物為食(shi),不會跟特(te)暴(bao)龍(long)(long)(long)(long)競(jing)爭食(shi)物。
在2001年,布魯斯·羅斯柴爾德(Bruce Rothschild)等人(ren)發表(biao)一(yi)份獸(shou)腳(jiao)類恐(kong)龍(long)的(de)壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe)與肌腱撕(si)裂傷研(yan)究,并研(yan)究它們的(de)行(xing)為模式。壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe)較常導因于習慣性(xing)動作,較少來自于外力(li)沖擊。他們研(yan)究18個(ge)(ge)特(te)暴龍(long)的(de)腳(jiao)掌(zhang)(zhang)骨(gu)(gu)(gu)骼,沒(mei)有發現(xian)壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe)的(de)跡(ji)象(xiang);而在10個(ge)(ge)被研(yan)究的(de)手(shou)(shou)掌(zhang)(zhang)骨(gu)(gu)(gu)頭中,其中一(yi)個(ge)(ge)手(shou)(shou)部(bu)骨(gu)(gu)(gu)頭有發現(xian)壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe)的(de)跡(ji)象(xiang)。腳(jiao)部(bu)骨(gu)(gu)(gu)頭的(de)壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe),可以歸咎(jiu)于奔跑(pao)、長(chang)途遷(qian)徙。手(shou)(shou)部(bu)骨(gu)(gu)(gu)頭的(de)壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe),極(ji)可能導因于捕抓(zhua)獵(lie)物而受傷。這些壓力(li)性(xing)骨(gu)(gu)(gu)折(zhe)、肌腱撕(si)裂傷跡(ji)象(xiang),顯示特(te)暴龍(long)常因為捕抓(zhua)獵(lie)物而受傷,而非吞(tun)食尸體。
在1946年,一(yi)個蘇聯(lian)與(yu)蒙古(gu)挖掘團隊在蒙古(gu)南戈壁省(sheng)的耐(nai)梅蓋特組發現一(yi)個接近(jin)完(wan)整的絕大(da)部(bu)分大(da)型頭顱骨與(yu)一(yi)些脊(ji)椎(zhui)骨,屬于(yu)一(yi)個大(da)型的獸腳類恐龍(long)。
在1955年,蘇聯(lian)古(gu)生物學家(jia)葉甫根尼(ni)·馬(ma)列夫(Evgeny Maleev)將這正模(mo)標本(PIN 551-1)建立為(wei)暴龍(long)的一個種,勇士(shi)暴龍(long)(Tyrannosaurus bataar)。種名(ming)баатар/baatar在蒙(meng)古(gu)語(yu)中意為(wei)"勇士(shi)",但被誤拼(pin)為(wei)bataar。
同年,馬列夫(fu)將3個(ge)獸(shou)腳類的化石敘述(shu)并命(ming)名,三者(zhe)都是頭顱(lu)骨,外加部分身(shen)體,全是同一個(ge)挖掘團隊在(zai)1948到49年間發(fa)(fa)現(xian)的。第一個(ge)(編號(hao)PIN 551-2)被(bei)建立為新(xin)屬,埃夫(fu)雷(lei)(lei)莫夫(fu)特暴龍(long)(T. efremovi),ταρβο?/tarbos在(zai)希(xi)臘文(wen)文(wen)中意(yi)為"恐怖(bu)的"、"敬(jing)畏的",而σαυρο?/saurus意(yi)為"蜥(xi)蜴",種名是以蘇聯古生物學家兼科幻小(xiao)說(shuo)作者(zhe)伊凡·埃夫(fu)雷(lei)(lei)莫夫(fu)(Ivan Yefremov)為名。另(ling)外兩個(ge)(編號(hao)PIN 553-1與PIN 552-2)被(bei)歸(gui)類于北美(mei)洲的蛇發(fa)(fa)女怪龍(long),G. lancinator與G. novojilovi。這三個(ge)標本都小(xiao)于1946年發(fa)(fa)現(xian)的標本。
自從蘇聯與(yu)蒙(meng)古的(de)挖掘團隊(dui)在40年代的(de)挖掘過(guo)后,一個波蘭(lan)與(yu)蒙(meng)古的(de)挖掘團隊(dui)再度(du)回到戈壁沙(sha)漠挖掘,從1963年持續(xu)到1971年,發(fa)現了許多新的(de)化(hua)石,并在耐(nai)梅蓋特(te)組發(fa)現了特(te)暴(bao)龍(long)的(de)新標本。
在1965年,蘇聯古生物(wu)學家(jia)阿納托利·康斯(si)坦丁諾維奇·羅(luo)特杰斯(si)特文斯(si)基(ji)(Anatoly Konstantinovich Rozhdestvensky)認為馬列夫所發(fa)現的標本(ben),其實(shi)是同種動物(wu)的不同生長階(jie)段,而且不同于北美洲的暴龍。
1979年(nian),董枝明在(zai)根(gen)據(ju)欒(luan)(luan)(luan)川縣(xian)發掘(jue)到五顆大(da)型牙(ya)齒命名為巒川暴(bao)龍(long)(Tyrannosaurus luanchuanensis)和霸王龍(long)相似(si)種(Tyrannosaurus cf. T. rex )的動物(wu)后來(lai)就被改為欒(luan)(luan)(luan)川特暴(bao)龍(long)等,而欒(luan)(luan)(luan)川特暴(bao)龍(long)實際也不能成立,因為后來(lai)的研究(jiu)把它(ta)歸入(ru)了勇士特暴(bao)龍(long),認(ren)為它(ta)實際只是(shi)勇士特暴(bao)龍(long)的青(qing)年(nian)個體。在(zai)欒(luan)(luan)(luan)川特暴(bao)龍(long)發現(xian)之前,廣東(dong)河源(yuan)發現(xian)過特暴(bao)龍(long)未定種的化石(shi)。發現(xian)欒(luan)(luan)(luan)川特暴(bao)龍(long)的秋扒(ba)組巖石(shi),秋扒(ba)組巖石(shi)是(shi)分(fen)布在(zai)潭(tan)頭盆地欒(luan)(luan)(luan)川縣(xian)一帶的白(bai)堊紀晚(wan)期地層,是(shi)由紫紅色的砂泥(ni)巖構成,最底部(bu)含有磚紅色的礫石(shi),其他發現(xian)過特暴(bao)龍(long)的地層也都屬于白(bai)堊紀晚(wan)期。
在1986年,雖(sui)然已(yi)經發(fa)現(xian)許多(duo)特(te)暴龍(long)(long)標本(ben),但很少已(yi)確定的(de)明確資(zi)料,特(te)暴龍(long)(long)被推論與(yu)其他暴龍(long)(long)科恐龍(long)(long)有(you)許多(duo)共同(tong)特(te)征。這些相近的(de)特(te)征讓(rang)某些科學家(jia)提(ti)出當時(shi)的(de)北美洲與(yu)歐(ou)亞大(da)陸(lu)之間(jian)有(you)可能(neng)的(de)連結,也(ye)許是(shi)陸(lu)橋(qiao)。他將四(si)個(ge)標本(ben)與(yu)新發(fa)現(xian)的(de)化(hua)石(shi),統(tong)合為勇士(shi)特(te)暴龍(long)(long)(T.bataar)。后來的(de)研究人員(yuan)都同(tong)意(yi)羅(luo)特(te)杰斯特(te)文斯基的(de)分類,包含馬(ma)列夫(fu)在內,但少數(shu)科學家(jia)使用埃夫(fu)雷莫夫(fu)特(te)暴龍(long)(long),而非勇士(shi)特(te)暴龍(long)(long)。
在(zai)1992年,美(mei)國古生(sheng)物學(xue)家肯尼思·卡彭(peng)特(Kenneth Carpenter)重新檢驗這些化石(shi),他根據頭顱骨的相似處,提出這些化石(shi)大部分屬(shu)于暴龍(long),并將勇士(shi)特暴龍(long)改回勇士(shi)暴龍(long);而蛇發女(nv)怪(guai)龍(long)的G. novojilovi是個較(jiao)小型的暴龍(long)科動物,另外建立為(wei)馬列(lie)夫龍(long)(Maleevosaurus novojilovi)。
在1995年,喬(qiao)治·奧利舍夫(fu)(fu)(fu)斯基(George Olshevsky)建(jian)立(li)了(le)勇(yong)士成(cheng)吉(ji)思(si)汗(han)龍(Jenghizkhan bataar),取(qu)代(dai)勇(yong)士特暴龍,并以成(cheng)吉(ji)思(si)汗(han)為名;他并提出(chu)埃夫(fu)(fu)(fu)雷莫夫(fu)(fu)(fu)特暴龍、馬列夫(fu)(fu)(fu)龍、勇(yong)士成(cheng)吉(ji)思(si)汗(han)龍是(shi)三(san)個(ge)獨立(li)的(de)屬,都(dou)生存于相(xiang)同時期(qi)的(de)耐梅蓋特組。
在1999年,一個研究提出馬列夫龍是特暴(bao)(bao)龍的幼年個體。自從1999年以(yi)來,所有的研究認為這些化石皆為同種動物,勇士特暴(bao)(bao)龍或是勇士暴(bao)(bao)龍。
在(zai)1993年到1998年,日本與蒙古的(de)挖掘(jue)團(tuan)隊,以及21世紀(ji)初,加拿大古生物學家菲(fei)力(li)·柯爾(er)(Phil Currie)的(de)挖掘(jue)小組,也發(fa)現(xian)了(le)許多特暴龍化石(shi)。不算上私(si)人(ren)收藏的(de)標(biao)本,已發(fa)現(xian)了(le)至少37個標(biao)本,包(bao)含超過15個完(wan)整(zheng)和部分(fen)的(de)頭顱(lu)骨化石(shi),與至少5個完(wan)整(zheng)的(de)顱(lu)后骨骸。
21世紀初,廣(guang)東河源盆地也出(chu)土過大(da)(da)型獸腳(jiao)類(lei)的大(da)(da)型暴龍類(lei)牙齒化石,推測(ce)可能是特暴龍。
特暴龍(long)的(de)(de)大部(bu)分化石,是(shi)成年(nian)或亞成年(nian)個(ge)(ge)(ge)(ge)體(ti),很(hen)少發現幼(you)年(nian)個(ge)(ge)(ge)(ge)體(ti)的(de)(de)化石。在2006年(nian)發現的(de)(de)一個(ge)(ge)(ge)(ge)幼(you)年(nian)個(ge)(ge)(ge)(ge)體(ti)身體(ti)骨(gu)骼(ge),帶有完(wan)整的(de)(de)頭(tou)顱骨(gu),頭(tou)部(bu)長度為29厘米,使得古生物學家可(ke)以(yi)研究特暴龍(long)的(de)(de)生長模式。這(zhe)(zhe)個(ge)(ge)(ge)(ge)幼(you)年(nian)個(ge)(ge)(ge)(ge)體(ti)化石,死亡時(shi)的(de)(de)年(nian)齡大約是(shi)2到3歲(sui)。與成年(nian)個(ge)(ge)(ge)(ge)體(ti)相(xiang)比,這(zhe)(zhe)個(ge)(ge)(ge)(ge)幼(you)年(nian)頭(tou)顱骨(gu)的(de)(de)結構(gou)虛(xu)弱,牙齒較細,顯(xian)示特暴龍(long)的(de)(de)幼(you)年(nian)個(ge)(ge)(ge)(ge)體(ti)、成年(nian)個(ge)(ge)(ge)(ge)體(ti)占據(ju)不同(tong)的(de)(de)生態位(wei),以(yi)免競爭相(xiang)同(tong)食物來源(yuan)。