特(te)暴(bao)龍(long)是發(fa)(fa)現于(yu)亞洲地區的(de)(de)(de)(de)(de)(de)(de)(de)大(da)型暴(bao)龍(long)類(lei),在外形上與北美(mei)洲的(de)(de)(de)(de)(de)(de)(de)(de)霸(ba)王龍(long)非常相似(si),是當(dang)時生態系統中的(de)(de)(de)(de)(de)(de)(de)(de)頂級捕(bu)食(shi)者。特(te)暴(bao)龍(long)的(de)(de)(de)(de)(de)(de)(de)(de)體型比霸(ba)王龍(long)和(he)諸城暴(bao)龍(long)略小,但(dan)(dan)明顯大(da)于(yu)其它的(de)(de)(de)(de)(de)(de)(de)(de)暴(bao)龍(long)類(lei),其頭(tou)骨(gu)的(de)(de)(de)(de)(de)(de)(de)(de)最大(da)長度可達1.3米(mi)。特(te)暴(bao)龍(long)的(de)(de)(de)(de)(de)(de)(de)(de)化石主要發(fa)(fa)現于(yu)蒙古(gu),但(dan)(dan)是也(ye)有一(yi)(yi)(yi)些證據表(biao)明這種(zhong)(zhong)巨型掠(lve)食(shi)者在中國的(de)(de)(de)(de)(de)(de)(de)(de)內蒙古(gu)以及新疆等(deng)地區都(dou)有分布(bu)。如同(tong)它在北美(mei)洲的(de)(de)(de)(de)(de)(de)(de)(de)近親一(yi)(yi)(yi)樣,特(te)暴(bao)龍(long)的(de)(de)(de)(de)(de)(de)(de)(de)標本量也(ye)非常巨大(da)。截止(zhi)到目(mu)前為(wei)止(zhi),古(gu)生物(wu)學(xue)家至少已經找(zhao)到了約30個特(te)暴(bao)龍(long)個體的(de)(de)(de)(de)(de)(de)(de)(de)標本,這其中不(bu)乏保存精(jing)美(mei)的(de)(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu)化石。在亞洲恐龍(long)古(gu)生物(wu)學(xue)發(fa)(fa)展的(de)(de)(de)(de)(de)(de)(de)(de)早期(qi)階段,特(te)暴(bao)龍(long)不(bu)同(tong)發(fa)(fa)育(yu)階段的(de)(de)(de)(de)(de)(de)(de)(de)個體曾經被當(dang)成過(guo)不(bu)同(tong)的(de)(de)(de)(de)(de)(de)(de)(de)物(wu)種(zhong)(zhong),不(bu)過(guo)在近期(qi)的(de)(de)(de)(de)(de)(de)(de)(de)研究中這些錯誤(wu)都(dou)被一(yi)(yi)(yi)一(yi)(yi)(yi)糾正。
特暴龍(long)的(de)屬(shu)名(ming)意為(wei)“可(ke)怕(pa)的(de)蜥蜴”,種名(ming)意為(wei)“勇士(shi)”。特暴龍(long)的(de)正型標(biao)本(ben)最(zui)初并沒有被歸(gui)入到特暴龍(long)屬(shu),而是被歸(gui)入到暴龍(long)屬(shu),并命名(ming)為(wei)“Tyrannosaurus bataar”,另(ling)外(wai)三個(ge)早期(qi)的(de)歸(gui)入標(biao)本(ben)中(zhong)有兩個(ge)被歸(gui)入了蛇發(fa)(fa)女怪龍(long)屬(shu)。直到1965年蘇聯古生(sheng)物學(xue)家(jia)阿納托利·康斯坦丁諾維奇·羅(luo)特杰(jie)斯特文斯基(ji)才發(fa)(fa)現馬列夫命名(ming)的(de)這(zhe)4件標(biao)本(ben)是同一物種不同發(fa)(fa)育階(jie)段的(de)個(ge)體,隨后它們(men)才被并入勇士(shi)特暴龍(long)。
正型(xing)標本:PIN 551-1保存(cun)了一部分頭骨(gu)和頸椎。最早被(bei)歸(gui)入暴龍(long)屬,后歸(gui)入特暴龍(long)屬。
此外蘭平(ping)暴(bao)龍(long)(Tyrannosaurus lanpingensis),吐(tu)魯番暴(bao)龍(long)(Tyrannosaurus turpanensis),Albertosaurus periculosus目前都被(bei)(bei)認為是屬于勇士(shi)特(te)暴(bao)龍(long)的(de)(de)不同(tong)個體發育階段(duan)。除此之外,在蒙古發現的(de)(de)分支龍(long)曾(ceng)經也(ye)被(bei)(bei)懷疑過是特(te)暴(bao)龍(long)的(de)(de)幼年個體,不過近年來由于虔州龍(long)的(de)(de)發現,分支龍(long)類的(de)(de)有(you)效性已經被(bei)(bei)學(xue)術界所接受。
特(te)暴(bao)龍(long)(long)(long)的(de)(de)(de)頭骨(gu)(gu)(gu)側視與霸(ba)(ba)(ba)王龍(long)(long)(long)非(fei)常相似。兩者(zhe)都具有(you)(you)背腹向加深的(de)(de)(de)頭骨(gu)(gu)(gu),且具有(you)(you)粗(cu)壯的(de)(de)(de)下頜。從背視可以看出兩種恐龍(long)(long)(long)的(de)(de)(de)頭骨(gu)(gu)(gu)整體輪廓(kuo)的(de)(de)(de)差異(yi),霸(ba)(ba)(ba)王龍(long)(long)(long)的(de)(de)(de)后(hou)端(duan)向外(wai)(wai)(wai)側極度(du)的(de)(de)(de)擴(kuo)展(zhan),使得雙(shuang)眼視覺范(fan)圍具有(you)(you)較大的(de)(de)(de)重疊。雖然特(te)暴(bao)龍(long)(long)(long)的(de)(de)(de)頭骨(gu)(gu)(gu)后(hou)端(duan)也有(you)(you)一定程(cheng)度(du)的(de)(de)(de)向外(wai)(wai)(wai)擴(kuo)展(zhan),但是與霸(ba)(ba)(ba)王龍(long)(long)(long)有(you)(you)較大的(de)(de)(de)差距。同時(shi)特(te)暴(bao)龍(long)(long)(long)的(de)(de)(de)吻(wen)端(duan)明顯(xian)比霸(ba)(ba)(ba)王龍(long)(long)(long)更(geng)窄。霸(ba)(ba)(ba)王龍(long)(long)(long)的(de)(de)(de)前頜骨(gu)(gu)(gu)和上頜骨(gu)(gu)(gu)相接的(de)(de)(de)骨(gu)(gu)(gu)縫上有(you)(you)一個(ge)明顯(xian)的(de)(de)(de)開孔暴(bao)露在頭骨(gu)(gu)(gu)的(de)(de)(de)外(wai)(wai)(wai)側面(mian),而這(zhe)一開孔在特(te)暴(bao)龍(long)(long)(long)頭骨(gu)(gu)(gu)的(de)(de)(de)外(wai)(wai)(wai)側面(mian)幾乎沒有(you)(you)。
特(te)暴龍(long)(long)的(de)(de)(de)(de)(de)(de)上(shang)頜(he)骨(gu)(gu)(gu)升(sheng)突粗狀發達(da),霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)較為(wei)纖細。兩種恐龍(long)(long)的(de)(de)(de)(de)(de)(de)鼻(bi)骨(gu)(gu)(gu)腹外側緣都具有(you)與上(shang)頜(he)骨(gu)(gu)(gu)相關節的(de)(de)(de)(de)(de)(de)溝,并幾(ji)乎前后向(xiang)延伸至鼻(bi)骨(gu)(gu)(gu)的(de)(de)(de)(de)(de)(de)整個長度(du)。但特(te)暴龍(long)(long)的(de)(de)(de)(de)(de)(de)關節面具有(you)很多脊,使鼻(bi)骨(gu)(gu)(gu)與上(shang)頜(he)骨(gu)(gu)(gu)的(de)(de)(de)(de)(de)(de)關節更(geng)緊(jin)密,而霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)鼻(bi)骨(gu)(gu)(gu)上(shang)頜(he)骨(gu)(gu)(gu)關節更(geng)簡單。相反的(de)(de)(de)(de)(de)(de)是,霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)鼻(bi)骨(gu)(gu)(gu)與淚(lei)骨(gu)(gu)(gu)的(de)(de)(de)(de)(de)(de)關節十分(fen)復雜且非(fei)常緊(jin)密,而特(te)暴龍(long)(long)的(de)(de)(de)(de)(de)(de)鼻(bi)骨(gu)(gu)(gu)與淚(lei)骨(gu)(gu)(gu)的(de)(de)(de)(de)(de)(de)關節則較為(wei)簡單。此外,霸(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)鼻(bi)骨(gu)(gu)(gu)比特(te)暴龍(long)(long)的(de)(de)(de)(de)(de)(de)更(geng)加寬大(da)。
霸(ba)(ba)王(wang)(wang)龍(long)的(de)(de)淚骨(gu)(gu)(gu)背(bei)緣較(jiao)特(te)暴龍(long)有更明(ming)顯的(de)(de)膨大。特(te)暴龍(long)的(de)(de)淚骨(gu)(gu)(gu)降(jiang)支較(jiao)為中空(kong),而霸(ba)(ba)王(wang)(wang)龍(long)的(de)(de)淚骨(gu)(gu)(gu)降(jiang)支幾乎是實心的(de)(de)。特(te)暴龍(long)的(de)(de)淚骨(gu)(gu)(gu)前(qian)突(tu)(tu)(tu)與霸(ba)(ba)王(wang)(wang)龍(long)的(de)(de)相比更為細長(chang),且(qie)前(qian)突(tu)(tu)(tu)關節面沒(mei)有分支。霸(ba)(ba)王(wang)(wang)龍(long)的(de)(de)淚骨(gu)(gu)(gu)前(qian)突(tu)(tu)(tu)關節面具有明(ming)顯的(de)(de)分叉。淚骨(gu)(gu)(gu)前(qian)突(tu)(tu)(tu)的(de)(de)內(nei)側面中段具有一個容(rong)納血管的(de)(de)開孔,這一開孔在霸(ba)(ba)王(wang)(wang)龍(long)的(de)(de)淚骨(gu)(gu)(gu)上的(de)(de)位(wei)置更靠前(qian)。
鱗骨(gu)(gu)的(de)(de)(de)(de)降支(zhi)(zhi)(zhi)與(yu)(yu)方(fang)顴(quan)骨(gu)(gu)的(de)(de)(de)(de)上支(zhi)(zhi)(zhi)具有很(hen)(hen)大的(de)(de)(de)(de)接觸面(mian)積,與(yu)(yu)艾伯(bo)塔龍(long)(long)(long)(long)(long)和(he)蛇(she)發(fa)(fa)女怪龍(long)(long)(long)(long)(long)不同(tong),而(er)與(yu)(yu)霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)非(fei)常相(xiang)似。在(zai)腹(fu)側(ce)視(shi),特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)顴(quan)骨(gu)(gu)不如霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)那樣很(hen)(hen)明(ming)顯(xian)(xian)的(de)(de)(de)(de)外翻。特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)和(he)霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)顴(quan)骨(gu)(gu)升突外側(ce)面(mian)都具有一不甚明(ming)顯(xian)(xian)的(de)(de)(de)(de)凹(ao)陷(xian)(xian),但(dan)是霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)這(zhe)一凹(ao)陷(xian)(xian)明(ming)顯(xian)(xian)小于(yu)特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)。兩者(zhe)的(de)(de)(de)(de)顴(quan)骨(gu)(gu)后(hou)(hou)突都具有二分(fen)支(zhi)(zhi)(zhi)的(de)(de)(de)(de)結構,區別(bie)在(zai)于(yu)特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)顴(quan)骨(gu)(gu)后(hou)(hou)突腹(fu)側(ce)支(zhi)(zhi)(zhi)的(de)(de)(de)(de)厚度(du)不均勻(yun),其腹(fu)緣較厚,向背緣發(fa)(fa)展(zhan)的(de)(de)(de)(de)過程中(zhong)逐(zhu)漸變薄(bo),而(er)霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)顴(quan)骨(gu)(gu)后(hou)(hou)突腹(fu)側(ce)支(zhi)(zhi)(zhi)的(de)(de)(de)(de)厚度(du)更(geng)(geng)為均一,且外側(ce)面(mian)更(geng)(geng)為明(ming)顯(xian)(xian)的(de)(de)(de)(de)隆起(qi)。特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)方(fang)顴(quan)骨(gu)(gu)柄明(ming)顯(xian)(xian)比霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)更(geng)(geng)加纖弱。霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)方(fang)顴(quan)骨(gu)(gu)前(qian)突比特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)更(geng)(geng)為寬大,能夠(gou)覆蓋住(zhu)顴(quan)骨(gu)(gu)后(hou)(hou)突的(de)(de)(de)(de)大部分(fen)面(mian)積,且內側(ce)面(mian)具有一個(ge)很(hen)(hen)大的(de)(de)(de)(de)凹(ao)陷(xian)(xian)。特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)顎骨(gu)(gu)上的(de)(de)(de)(de)內鼻(bi)孔邊(bian)緣發(fa)(fa)育的(de)(de)(de)(de)很(hen)(hen)明(ming)顯(xian)(xian),而(er)這(zhe)一特(te)(te)征在(zai)霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)顎骨(gu)(gu)上相(xiang)對較弱。特(te)(te)暴(bao)(bao)龍(long)(long)(long)(long)(long)外翼骨(gu)(gu)上的(de)(de)(de)(de)氣孔比霸(ba)(ba)王(wang)(wang)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)更(geng)(geng)小。
特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)犁骨(gu)(gu)前(qian)突(tu)向(xiang)前(qian)極度延(yan)伸(shen),與前(qian)頜骨(gu)(gu)的(de)(de)(de)(de)(de)(de)顎突(tu)相接(jie),而(er)這一特(te)征在(zai)霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)中不(bu)明顯。特(te)暴(bao)龍(long)(long)和(he)(he)霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)翼骨(gu)(gu)的(de)(de)(de)(de)(de)(de)最大區(qu)別在(zai)于(yu)顎骨(gu)(gu)關節面的(de)(de)(de)(de)(de)(de)前(qian)邊(bian)緣。霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)翼骨(gu)(gu)額骨(gu)(gu)突(tu)的(de)(de)(de)(de)(de)(de)前(qian)邊(bian)緣是一條斜線,而(er)特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)翼骨(gu)(gu)顎骨(gu)(gu)突(tu)前(qian)邊(bian)緣向(xiang)前(qian)突(tu)出。特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)上(shang)(shang)枕(zhen)骨(gu)(gu)的(de)(de)(de)(de)(de)(de)內外向(xiang)寬度小于(yu)枕(zhen)髁,而(er)霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)上(shang)(shang)枕(zhen)骨(gu)(gu)更(geng)寬。從側面看,特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)齒骨(gu)(gu)吻端比霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)更(geng)加(jia)尖(jian)細一點。霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)夾(jia)板(ban)(ban)骨(gu)(gu)腹緣相較(jiao)于(yu)特(te)暴(bao)龍(long)(long)有更(geng)大的(de)(de)(de)(de)(de)(de)傾斜角度,并且夾(jia)板(ban)(ban)骨(gu)(gu)的(de)(de)(de)(de)(de)(de)前(qian)端位(wei)于(yu)更(geng)高的(de)(de)(de)(de)(de)(de)位(wei)置(zhi)。特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)上(shang)(shang)隅骨(gu)(gu)孔較(jiao)霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)的(de)(de)(de)(de)(de)(de)要(yao)發(fa)達。霸(ba)(ba)(ba)王(wang)(wang)龍(long)(long)和(he)(he)特(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)(de)(de)成(cheng)體(ti)上(shang)(shang)頜骨(gu)(gu)齒和(he)(he)齒骨(gu)(gu)齒的(de)(de)(de)(de)(de)(de)數量不(bu)一樣。
歸(gui)入(ru)標本:PIN 551-2是一具接近(jin)完整(zheng)的骨架。最(zui)早(zao)被歸(gui)入(ru)到(dao)特暴(bao)龍(long)屬(shu)(shu)的歸(gui)入(ru)種埃(ai)夫雷莫夫特暴(bao)龍(long)(Tarbosaurus efremovi)。現已(yi)歸(gui)入(ru)到(dao)勇(yong)(yong)士特暴(bao)龍(long),且(qie)勇(yong)(yong)士特暴(bao)龍(long)是特暴(bao)龍(long)屬(shu)(shu)下(xia)的唯一有效種。
歸入標本:PIN 553-1保存了頭骨(gu)(gu),背椎和(he)尾椎,掌骨(gu)(gu)和(he)跖骨(gu)(gu)。最(zui)開始被歸入到蛇發(fa)女怪龍屬。現已歸入勇士特暴龍。
歸入標本:PIN 552-2保存了頭(tou)骨和頭(tou)后骨骼。最開始(shi)被(bei)歸入到蛇發女怪龍屬。現已歸入勇士特暴龍。
歸入(ru)標本(ben)(ben):MPC-D 107/7是一件幼(you)年的標本(ben)(ben),關(guan)聯保(bao)(bao)存。除了缺少頸椎(zhui),前部(bu)背(bei)椎(zhui)和末(mo)端尾(wei)椎(zhui),其它骨頭基(ji)本(ben)(ben)都有保(bao)(bao)存。
歸入(ru)標本(ben):ZPAL MgD-I/3幾乎完(wan)整的(de)(de)(de)中等(deng)體(ti)型的(de)(de)(de)個體(ti)。保存(cun)了頭骨(gu),頸椎(zhui)和背椎(zhui),10節(jie)近(jin)端(duan)尾椎(zhui),完(wan)整的(de)(de)(de)腸(chang)骨(gu),恥骨(gu)和坐骨(gu),肋(lei)骨(gu)和腹膜肋(lei),完(wan)整的(de)(de)(de)肩胛骨(gu),烏喙(hui)骨(gu),左側的(de)(de)(de)前肢,不(bu)完(wan)整保存(cun)的(de)(de)(de)右側后肢。
歸入標本:ZPAL Mgd-I/4部(bu)分保(bao)存的大(da)型個(ge)體。保(bao)存了左側后肢(zhi),腸骨,13節椎體。
歸(gui)入標本:ZPAL Mgd-I/5不完整保(bao)存(cun)的(de)(de)大(da)型骨(gu)(gu)(gu)架。保(bao)存(cun)了左側(ce)(ce)(ce)上頜(he)骨(gu)(gu)(gu),左側(ce)(ce)(ce)方骨(gu)(gu)(gu),左側(ce)(ce)(ce)的(de)(de)下(xia)頜(he)和(he)(he)右側(ce)(ce)(ce)的(de)(de)下(xia)頜(he)碎(sui)(sui)片,11節左側(ce)(ce)(ce)肋骨(gu)(gu)(gu)的(de)(de)碎(sui)(sui)片,恥骨(gu)(gu)(gu)碎(sui)(sui)片,坐骨(gu)(gu)(gu),腸骨(gu)(gu)(gu)碎(sui)(sui)片,左側(ce)(ce)(ce)后(hou)肢(zhi)和(he)(he)右側(ce)(ce)(ce)跖骨(gu)(gu)(gu),和(he)(he)很(hen)多碎(sui)(sui)屑。
歸入標本:ZPAL Mgd-I/26不完整(zheng)保存(cun)的上頜骨(gu)和一部(bu)分(fen)保存(cun)下來的牙齒。
歸入標本:ZPAL Mgd-I/29不完(wan)(wan)(wan)整的大型骨架。保(bao)存了頭(tou)骨和(he)(he)較為(wei)完(wan)(wan)(wan)整的齒列。頭(tou)后骨骼保(bao)存了6節頸椎,5節薦椎,22節尾椎,11節右(you)側(ce)肋骨,腸(chang)骨,不完(wan)(wan)(wan)整的恥(chi)骨和(he)(he)近端(duan)坐骨,左側(ce)肱骨,尺骨和(he)(he)橈骨的遠端(duan),第(di)一手指,幾(ji)乎完(wan)(wan)(wan)整的右(you)側(ce)后肢。
歸入標本:ZPAL Mgd-I/31右側上(shang)頜骨的近端。
歸入標本(ben):ZPAL Mgd-I/34頭骨右側(ce)的碎片(pian)。
歸入標(biao)本:ZPAL Mgd-I/38不完全保存的頭骨(gu),12節肋骨(gu),右(you)(you)側(ce)(ce)股骨(gu)的遠端(duan)。右(you)(you)側(ce)(ce)脛骨(gu)的遠端(duan),右(you)(you)側(ce)(ce)的3-4跖(zhi)骨(gu),第4趾(zhi)的近端(duan)趾(zhi)節骨(gu)。
歸入標(biao)本:ZPAL Mgd-I/44頭骨的(de)右側。
歸入標本:ZPAL Mgd-I/45頭骨(gu)碎片(pian)。
歸入(ru)標本:ZPAL Mgd-I/46右側下頜碎片和其它(ta)7塊頭骨碎片,和兩節斷掉的背椎肋骨。
歸入標本:ZPAL Mgd-I/52左側下頜齒骨。
歸入標本:ZPAL Mgd-I/67右側顴骨。
歸入標本:ZPAL Mgd-I/93頭(tou)骨腦顱(lu)。
歸入(ru)標本(ben):ZPAL Mgd-I/109大的帶有圍巖的頭骨。
歸入標本:ZPAL Mgd-I/178頭(tou)骨(gu)的碎片,脊(ji)椎骨(gu)和股骨(gu)。
歸入(ru)標本:GIN 100/61頭骨碎片和(he)頭后骨骼。
歸入標本:GIN 100/62頭骨(gu)碎片和頭后(hou)骨(gu)骼。
歸入標本:GIN 100/65頭骨的右半部分。
歸入標本(ben):GIN 100/67頭骨的碎(sui)片,腦(nao)顱。
歸(gui)入標(biao)本:GIN 100/69枕區。
歸入標本:GIN 100/70頭骨碎(sui)片和椎體(ti)。
歸入標本:GIN 107/2完整的(de)骨(gu)架。
歸入標(biao)本(ben):GIN 107/3頭骨。
歸(gui)入標本(ben):PIN 551-3骨架(jia),保存狀況未描述。最(zui)早被歸(gui)入到(dao)特(te)暴龍屬的歸(gui)入種埃夫雷(lei)莫夫特(te)暴龍。現(xian)已歸(gui)入到(dao)勇士(shi)特(te)暴龍。
歸入(ru)標本(ben):PIN 551-4骨架,保存狀況(kuang)未描(miao)述。最(zui)早被歸入(ru)到(dao)特暴(bao)龍(long)屬的歸入(ru)種埃夫(fu)雷莫夫(fu)特暴(bao)龍(long)。現已歸入(ru)到(dao)勇士特暴(bao)龍(long)。
歸(gui)(gui)(gui)入(ru)(ru)(ru)標本(ben):PIN 551-91右側上頜(he)骨。最早被歸(gui)(gui)(gui)入(ru)(ru)(ru)到特暴龍屬的歸(gui)(gui)(gui)入(ru)(ru)(ru)種埃夫雷莫夫特暴龍。現已歸(gui)(gui)(gui)入(ru)(ru)(ru)到勇士特暴龍。
歸(gui)入(ru)標本:PIN 552-1目前(qian)僅有復制品(pin)保存。最早(zao)被歸(gui)入(ru)到特(te)暴龍(long)(long)屬(shu)的(de)歸(gui)入(ru)種埃夫(fu)雷莫夫(fu)特(te)暴龍(long)(long)。現已(yi)歸(gui)入(ru)到勇士(shi)特(te)暴龍(long)(long)。
歸入(ru)標本(ben):PIN 553-2保存狀況未描(miao)述。早被歸入(ru)到特暴龍(long)屬的歸入(ru)種埃夫雷莫(mo)夫特暴龍(long)。現已歸入(ru)到勇士特暴龍(long)。
歸入標本:IVPP V 4878火焰(yan)山鄯善龍(Shanshanosaurus huoyanshanensis)的(de)正(zheng)型標本,發現于中國新疆,現在認為可能是特暴(bao)龍的(de)幼體(ti)。保(bao)存了上頜骨(gu),下頜大部分骨(gu)塊,多節(jie)頸椎(zhui),背椎(zhui)和(he)薦椎(zhui),肩(jian)帶(dai)骨(gu)骼(ge),肱骨(gu),恥骨(gu)遠端(duan),股骨(gu)和(he)脛骨(gu)近(jin)端(duan)。
歸入(ru)標本:IVPP V 4733欒川暴(bao)龍(long)(Tyrannosaurus luanchuanensis)的正型標本,為5顆(ke)牙齒,現認為也(ye)可(ke)能(neng)屬于特暴(bao)龍(long)。
歸入標本:IVPP V 836為(wei)破碎金剛口龍(Chingkankousaurus fragilis)的(de)(de)正型標本,為(wei)一節不完整的(de)(de)肩胛骨(gu)標本。布魯薩特(te)等人認為(wei)很有(you)可能是特(te)暴龍,但不太(tai)確(que)定。
特暴(bao)(bao)(bao)龍(long)(long)同霸(ba)王龍(long)(long)一樣,是暴(bao)(bao)(bao)龍(long)(long)超科中(zhong)最進步的(de)成員。大多數的(de)系統發育分析都支持霸(ba)王龍(long)(long)和(he)特暴(bao)(bao)(bao)龍(long)(long)構成一個(ge)姐妹群,諸(zhu)城(cheng)暴(bao)(bao)(bao)龍(long)(long)作為兩者所構成單(dan)系的(de)姐妹群。也有一些分析認為特暴(bao)(bao)(bao)龍(long)(long)與諸(zhu)城(cheng)暴(bao)(bao)(bao)龍(long)(long)的(de)親(qin)緣關系更近。
如果不考慮標本(ben)量很(hen)少且質量很(hen)差的(de)(de)(de)諸城暴(bao)龍(long),特(te)暴(bao)龍(long)的(de)(de)(de)體型基本(ben)穩坐暴(bao)龍(long)類(lei)的(de)(de)(de)第二把(ba)交椅。最大的(de)(de)(de)特(te)暴(bao)龍(long)頭骨長(chang)度達到1.3米,這一大小顯著的(de)(de)(de)大于其它的(de)(de)(de)大型暴(bao)龍(long)類(lei),并(bing)基本(ben)達到了(le)個別霸(ba)王龍(long)成(cheng)年標本(ben)的(de)(de)(de)頭骨長(chang)度。目前最大的(de)(de)(de)特(te)暴(bao)龍(long)標本(ben)體長(chang)是肯(ken)定能(neng)超過10米的(de)(de)(de)。特(te)暴(bao)龍(long)的(de)(de)(de)體重(zhong)一般(ban)估計(ji)在4噸左(zuo)右(you)。
通(tong)過(guo)(guo)(guo)(guo)仔細對比霸(ba)王(wang)龍(long)及(ji)其它(ta)北(bei)(bei)美(mei)暴(bao)(bao)(bao)(bao)龍(long)亞科成員的(de)(de)(de)頭(tou)(tou)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)各個骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)塊骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)縫與(yu)特暴(bao)(bao)(bao)(bao)龍(long)的(de)(de)(de)差異(yi)(yi)(yi)可(ke)以發現霸(ba)王(wang)龍(long)與(yu)特暴(bao)(bao)(bao)(bao)龍(long)的(de)(de)(de)頭(tou)(tou)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)具(ju)有(you)不(bu)太一樣的(de)(de)(de)生(sheng)物力(li)學(xue)結(jie)構。大型暴(bao)(bao)(bao)(bao)龍(long)類都(dou)具(ju)有(you)毀滅性的(de)(de)(de)咬(yao)合力(li),而它(ta)們的(de)(de)(de)頭(tou)(tou)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)也(ye)(ye)必(bi)須(xu)在(zai)撕(si)咬(yao)獵物的(de)(de)(de)同(tong)時(shi)承受同(tong)樣巨大的(de)(de)(de)反作用力(li)。北(bei)(bei)美(mei)洲的(de)(de)(de)暴(bao)(bao)(bao)(bao)龍(long)亞科主要通(tong)過(guo)(guo)(guo)(guo)上頜骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)-鼻(bi)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)-淚(lei)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)-額骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)/前額骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)的(de)(de)(de)緊密(mi)關(guan)(guan)(guan)節(jie)(jie)(jie)構成來承受撕(si)咬(yao)獵物過(guo)(guo)(guo)(guo)程中(zhong)產生(sheng)的(de)(de)(de)反作用力(li)。其中(zhong),鼻(bi)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)與(yu)淚(lei)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)構成了非(fei)常(chang)(chang)強(qiang)的(de)(de)(de)關(guan)(guan)(guan)節(jie)(jie)(jie),且這一特征普遍(bian)存(cun)在(zai)于(yu)(yu)北(bei)(bei)美(mei)的(de)(de)(de)大型暴(bao)(bao)(bao)(bao)龍(long)亞科中(zhong)。對于(yu)(yu)特暴(bao)(bao)(bao)(bao)龍(long)的(de)(de)(de)研究顯(xian)示,與(yu)北(bei)(bei)美(mei)洲的(de)(de)(de)親(qin)戚們相比,特暴(bao)(bao)(bao)(bao)龍(long)的(de)(de)(de)淚(lei)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)前突(tu)向下(xia)偏轉的(de)(de)(de)更多,并且擁(yong)有(you)比北(bei)(bei)美(mei)洲暴(bao)(bao)(bao)(bao)龍(long)類更為強(qiang)壯的(de)(de)(de)上頜骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)升突(tu),使得(de)這兩個塊骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)頭(tou)(tou)通(tong)過(guo)(guo)(guo)(guo)復雜的(de)(de)(de)關(guan)(guan)(guan)節(jie)(jie)(jie)面緊密(mi)關(guan)(guan)(guan)節(jie)(jie)(jie)在(zai)一起(qi)。同(tong)時(shi)特暴(bao)(bao)(bao)(bao)龍(long)的(de)(de)(de)前額骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)與(yu)淚(lei)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)也(ye)(ye)有(you)非(fei)常(chang)(chang)緊密(mi)的(de)(de)(de)關(guan)(guan)(guan)節(jie)(jie)(jie)。非(fei)常(chang)(chang)強(qiang)大的(de)(de)(de)上頜骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)-淚(lei)骨(gu)(gu)(gu)(gu)(gu)(gu)(gu)關(guan)(guan)(guan)節(jie)(jie)(jie)也(ye)(ye)見于(yu)(yu)異(yi)(yi)(yi)特龍(long)和中(zhong)華(hua)盜(dao)龍(long)這樣的(de)(de)(de)大型異(yi)(yi)(yi)特龍(long)類中(zhong)。
對(dui)于異特(te)(te)龍(long)(long)(long)(long)(long)頭骨(gu)的(de)(de)(de)(de)(de)(de)(de)有限元分析(xi)顯示(shi),異特(te)(te)龍(long)(long)(long)(long)(long)在(zai)攻擊(ji)獵物時,頭骨(gu)的(de)(de)(de)(de)(de)(de)(de)上(shang)頜(he)骨(gu)-淚骨(gu)關(guan)節處會承受(shou)非(fei)(fei)常(chang)大(da)的(de)(de)(de)(de)(de)(de)(de)反作用(yong)力(li)(li)。另外一(yi)點區(qu)別(bie)是,北美(mei)洲的(de)(de)(de)(de)(de)(de)(de)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)下頜(he)的(de)(de)(de)(de)(de)(de)(de)齒骨(gu)和(he)后面的(de)(de)(de)(de)(de)(de)(de)骨(gu)塊具有較(jiao)為靈活的(de)(de)(de)(de)(de)(de)(de)關(guan)節,使得它(ta)們的(de)(de)(de)(de)(de)(de)(de)下頜(he)可以(yi)一(yi)定(ding)程度(du)的(de)(de)(de)(de)(de)(de)(de)內外擴展和(he)收縮,而特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)沒有這樣(yang)的(de)(de)(de)(de)(de)(de)(de)關(guan)節,使得它(ta)們的(de)(de)(de)(de)(de)(de)(de)下頜(he)更為一(yi)體化。造成這些區(qu)別(bie)的(de)(de)(de)(de)(de)(de)(de)原因主(zhu)要(yao)是亞洲暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)和(he)北美(mei)洲的(de)(de)(de)(de)(de)(de)(de)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)所(suo)面對(dui)的(de)(de)(de)(de)(de)(de)(de)獵物不太(tai)一(yi)樣(yang)。晚白堊世(shi)北美(mei)洲的(de)(de)(de)(de)(de)(de)(de)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)主(zhu)要(yao)捕(bu)食角龍(long)(long)(long)(long)(long)類(lei)和(he)鴨嘴龍(long)(long)(long)(long)(long)類(lei),而亞洲的(de)(de)(de)(de)(de)(de)(de)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)則有更多的(de)(de)(de)(de)(de)(de)(de)機會捕(bu)獵大(da)型(xing)(xing)蜥腳類(lei)恐龍(long)(long)(long)(long)(long),比如泰坦巨龍(long)(long)(long)(long)(long)類(lei)。目前還未有研究使用(yong)模型(xing)(xing)化的(de)(de)(de)(de)(de)(de)(de)方法精確計(ji)算過特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)(de)(de)(de)咬合力(li)(li),但定(ding)性的(de)(de)(de)(de)(de)(de)(de)比較(jiao)認為,特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)與其它(ta)大(da)型(xing)(xing)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)一(yi)樣(yang)擁有非(fei)(fei)常(chang)巨大(da)的(de)(de)(de)(de)(de)(de)(de)咬合力(li)(li)。
菲(fei)利普.柯瑞等人(ren)在2003年的(de)(de)(de)一(yi)項研究中報(bao)道了(le)2個(ge)可能屬于特(te)暴龍(long)的(de)(de)(de)腳印(yin)。這些(xie)腳印(yin)非常巨大(da),有(you)61厘米長,保存了(le)第三趾爪在地(di)上留下的(de)(de)(de)痕(hen)跡。同時放大(da)觀察腳印(yin)還可以看(kan)到里面的(de)(de)(de)皮(pi)(pi)膚印(yin)痕(hen)。皮(pi)(pi)膚印(yin)痕(hen)中的(de)(de)(de)每個(ge)鱗片大(da)約(yue)有(you)2毫米寬(kuan)。另外一(yi)件(jian)特(te)暴龍(long)的(de)(de)(de)骨架上也發現了(le)皮(pi)(pi)膚印(yin)痕(hen)。皮(pi)(pi)膚印(yin)痕(hen)保留在接近咽(yan)喉的(de)(de)(de)位置,鱗片的(de)(de)(de)寬(kuan)度約(yue)為2.4毫米。遺憾的(de)(de)(de)是這件(jian)標本已經(jing)被破(po)壞(huai),無法對其進行更(geng)深入的(de)(de)(de)研究。
一件(jian)特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)標本保存(cun)了完(wan)整(zheng)的(de)(de)(de)(de)(de)(de)顱腔。通過制作顱內模(mo)可(ke)以了解特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)腦(nao)結構。特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)腦(nao)結構與霸(ba)王(wang)龍(long)(long)(long)十分相似,其主(zhu)要的(de)(de)(de)(de)(de)(de)區別(bie)僅存(cun)在(zai)于(yu)個別(bie)腦(nao)神(shen)經(jing)基(ji)部(bu)的(de)(de)(de)(de)(de)(de)位置,如(ru)(ru)(ru)三叉神(shen)經(jing)和副(fu)神(shen)經(jing)。一只12米(mi)長的(de)(de)(de)(de)(de)(de)特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)腦(nao)體(ti)積約(yue)為184立方厘米(mi)。特(te)(te)暴(bao)(bao)龍(long)(long)(long)和霸(ba)王(wang)龍(long)(long)(long)一樣擁(yong)有(you)(you)非常大(da)的(de)(de)(de)(de)(de)(de)嗅葉,這說明(ming)(ming)它(ta)(ta)(ta)們(men)有(you)(you)良好(hao)的(de)(de)(de)(de)(de)(de)嗅覺(jue)。同(tong)時它(ta)(ta)(ta)們(men)也(ye)(ye)具有(you)(you)發(fa)(fa)達的(de)(de)(de)(de)(de)(de)犁(li)鼻器用于(yu)探測荷(he)爾蒙(meng),這說明(ming)(ming)特(te)(te)暴(bao)(bao)龍(long)(long)(long)可(ke)能擁(yong)有(you)(you)復雜的(de)(de)(de)(de)(de)(de)交(jiao)配行為。特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)聽(ting)神(shen)經(jing)也(ye)(ye)很發(fa)(fa)達,說明(ming)(ming)它(ta)(ta)(ta)們(men)有(you)(you)較好(hao)的(de)(de)(de)(de)(de)(de)聽(ting)力。特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)中腦(nao)頂部(bu)(midbrain tectum)不甚發(fa)(fa)達,它(ta)(ta)(ta)的(de)(de)(de)(de)(de)(de)動眼(yan)神(shen)經(jing)和視神(shen)經(jing)也(ye)(ye)是如(ru)(ru)(ru)此(ci)。在(zai)頭骨(gu)的(de)(de)(de)(de)(de)(de)宏觀形(xing)態(tai)形(xing)態(tai)上,特(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)(de)(de)頭骨(gu)后(hou)端向外側(ce)擴展的(de)(de)(de)(de)(de)(de)較少,其雙(shuang)眼(yan)視覺(jue)疊加范圍不如(ru)(ru)(ru)霸(ba)王(wang)龍(long)(long)(long)。因此(ci)特(te)(te)暴(bao)(bao)龍(long)(long)(long)在(zai)生活的(de)(de)(de)(de)(de)(de)時候對(dui)嗅覺(jue)和聽(ting)覺(jue)的(de)(de)(de)(de)(de)(de)依賴要遠大(da)于(yu)對(dui)視覺(jue)的(de)(de)(de)(de)(de)(de)依賴。
同北(bei)(bei)美的(de)(de)(de)(de)(de)(de)大型暴(bao)(bao)龍(long)類一樣,特(te)暴(bao)(bao)龍(long)在(zai)生長(chang)發(fa)(fa)育過程中也伴隨著一系列的(de)(de)(de)(de)(de)(de)形態變化。目前發(fa)(fa)現(xian)的(de)(de)(de)(de)(de)(de)大多數(shu)特(te)暴(bao)(bao)龍(long)標本(ben)屬于亞(ya)成(cheng)年(nian)(nian)(nian)個(ge)體(ti)或成(cheng)年(nian)(nian)(nian)個(ge)體(ti),只(zhi)有(you)(you)很少的(de)(de)(de)(de)(de)(de)幼(you)(you)(you)(you)年(nian)(nian)(nian)個(ge)體(ti)。2011年(nian)(nian)(nian)發(fa)(fa)表(biao)的(de)(de)(de)(de)(de)(de)一件(jian)(jian)特(te)暴(bao)(bao)龍(long)幼(you)(you)(you)(you)體(ti)標本(ben)使得古(gu)生物學家(jia)對于特(te)暴(bao)(bao)龍(long)的(de)(de)(de)(de)(de)(de)幼(you)(you)(you)(you)年(nian)(nian)(nian)時期(qi)有(you)(you)了更多了解(jie)。這只(zhi)幼(you)(you)(you)(you)體(ti)在(zai)死亡的(de)(de)(de)(de)(de)(de)時候(hou)大約只(zhi)有(you)(you)2-3歲。與成(cheng)年(nian)(nian)(nian)個(ge)體(ti)相比,幼(you)(you)(you)(you)體(ti)的(de)(de)(de)(de)(de)(de)頭骨較為脆(cui)弱(ruo),牙齒(chi)也不夠強壯(zhuang),說明幼(you)(you)(you)(you)體(ti)的(de)(de)(de)(de)(de)(de)獵物與成(cheng)體(ti)有(you)(you)較大的(de)(de)(de)(de)(de)(de)不同。與霸王(wang)龍(long)和(he)懼龍(long)不同的(de)(de)(de)(de)(de)(de)是(shi),這件(jian)(jian)幼(you)(you)(you)(you)年(nian)(nian)(nian)特(te)暴(bao)(bao)龍(long)標本(ben)的(de)(de)(de)(de)(de)(de)齒(chi)骨齒(chi)和(he)上(shang)頜(he)骨齒(chi)的(de)(de)(de)(de)(de)(de)數(shu)量與成(cheng)年(nian)(nian)(nian)個(ge)體(ti)的(de)(de)(de)(de)(de)(de)數(shu)量一致(zhi),說明特(te)暴(bao)(bao)龍(long)在(zai)個(ge)體(ti)發(fa)(fa)育的(de)(de)(de)(de)(de)(de)過程中不存(cun)在(zai)牙齒(chi)數(shu)量變化的(de)(de)(de)(de)(de)(de)現(xian)象。這一區(qu)別(bie)是(shi)否廣泛存(cun)在(zai)與亞(ya)洲和(he)北(bei)(bei)美洲的(de)(de)(de)(de)(de)(de)其它(ta)暴(bao)(bao)龍(long)亞(ya)科(ke)物種還需要進一步(bu)檢驗。對于幼(you)(you)(you)(you)體(ti)鞏膜環的(de)(de)(de)(de)(de)(de)研究表(biao)明,幼(you)(you)(you)(you)體(ti)可能具(ju)有(you)(you)夜行(xing)的(de)(de)(de)(de)(de)(de)習性。成(cheng)年(nian)(nian)(nian)特(te)暴(bao)(bao)龍(long)是(shi)否具(ju)有(you)(you)夜行(xing)習性還需要更多的(de)(de)(de)(de)(de)(de)化石(shi)證(zheng)據(ju)支(zhi)持。
特暴(bao)龍(long)(long)(long)(long)(long)是其(qi)所(suo)處生態(tai)系(xi)統中最(zui)大(da)的(de)(de)捕食者,與其(qi)共存的(de)(de)植食性恐龍(long)(long)(long)(long)(long)包(bao)括蜥腳(jiao)類的(de)(de)耐(nai)梅(mei)蓋(gai)特龍(long)(long)(long)(long)(long)和(he)(he)(he)后(hou)凹尾龍(long)(long)(long)(long)(long),甲龍(long)(long)(long)(long)(long)類的(de)(de)美甲龍(long)(long)(long)(long)(long),腫頭龍(long)(long)(long)(long)(long)類的(de)(de)傾頭龍(long)(long)(long)(long)(long),鴨(ya)嘴龍(long)(long)(long)(long)(long)類的(de)(de)櫛龍(long)(long)(long)(long)(long)和(he)(he)(he)巴思缽氏(shi)龍(long)(long)(long)(long)(long),屬(shu)于(yu)(yu)鐮(lian)刀龍(long)(long)(long)(long)(long)類的(de)(de)鐮(lian)刀龍(long)(long)(long)(long)(long),屬(shu)于(yu)(yu)似鳥龍(long)(long)(long)(long)(long)類似鵝(e)龍(long)(long)(long)(long)(long),似雞(ji)龍(long)(long)(long)(long)(long)和(he)(he)(he)體型巨大(da)的(de)(de)恐手龍(long)(long)(long)(long)(long)。肉食性恐龍(long)(long)(long)(long)(long)則包(bao)括同屬(shu)于(yu)(yu)暴(bao)龍(long)(long)(long)(long)(long)亞(ya)科(ke)的(de)(de)分(fen)支龍(long)(long)(long)(long)(long),屬(shu)于(yu)(yu)傷齒龍(long)(long)(long)(long)(long)類的(de)(de)無聊龍(long)(long)(long)(long)(long),鴕鳥龍(long)(long)(long)(long)(long)和(he)(he)(he)扎(zha)納巴扎(zha)爾龍(long)(long)(long)(long)(long),屬(shu)于(yu)(yu)竊蛋龍(long)(long)(long)(long)(long)類的(de)(de)單足龍(long)(long)(long)(long)(long),耐(nai)梅(mei)蓋(gai)特母(mu)龍(long)(long)(long)(long)(long)和(he)(he)(he)瑞欽(qin)龍(long)(long)(long)(long)(long)。
特(te)暴龍(long)曾登場于BBC的紀錄片《恐龍(long)兇面目》以及《與龍(long)同(tong)行特(te)輯(ji):尋(xun)爪記》。近(jin)期(qi)它又在紀錄片《史(shi)前星球(qiu)》中(zhong)登場。
在電影方面,特暴龍登場于韓國(guo)的動畫電影《韓半島的恐(kong)龍》以及國(guo)產恐(kong)龍動畫電影《恐(kong)龍王》。
特暴龍位于食物鏈的頂端,是(shi)一種(zhong)頂級掠(lve)食(shi)動物。特(te)暴(bao)(bao)龍(long)(long)(long)在暴(bao)(bao)龍(long)(long)(long)科(ke)(ke)中(zhong)的分類位置仍未確定。以前(qian),有(you)些(xie)(xie)科(ke)(ke)學家認為勇士特(te)暴(bao)(bao)龍(long)(long)(long)其實是(shi)北美洲(zhou)暴(bao)(bao)龍(long)(long)(long)的亞洲(zhou)種(zhong)。如(ru)果屬實,將使(shi)(shi)特(te)暴(bao)(bao)龍(long)(long)(long)成為無效的分類。即(ji)使(shi)(shi)特(te)暴(bao)(bao)龍(long)(long)(long)與暴(bao)(bao)龍(long)(long)(long)不是(shi)同種(zhong)動物,它(ta)們被認為有(you)接(jie)近(jin)的親緣關(guan)系。有(you)些(xie)(xie)科(ke)(ke)學家認為,同樣發現于(yu)蒙古(gu)的分支龍(long)(long)(long),是(shi)特(te)暴(bao)(bao)龍(long)(long)(long)的近(jin)親。
特暴龍的(de)(de)化石記(ji)錄(lu)保(bao)存良好,已有數十(shi)個標本,包含至少5個完(wan)整的(de)(de)頭顱(lu)骨與骨骸。這些化石讓科學家得以(yi)研究它(ta)們的(de)(de)種系發生學、頭部力(li)學、以(yi)及(ji)腦(nao)部結(jie)構(gou)。
特暴(bao)龍(long)(long)屬于暴(bao)龍(long)(long)科(ke)(ke)的(de)(de)暴(bao)龍(long)(long)亞(ya)(ya)科(ke)(ke)。該亞(ya)(ya)科(ke)(ke)還包(bao)含較(jiao)(jiao)早期(qi)(qi)的(de)(de)懼(ju)龍(long)(long)、較(jiao)(jiao)晚期(qi)(qi)的(de)(de)暴(bao)龍(long)(long),都發(fa)現于北美洲,可能還有蒙(meng)古的(de)(de)分支龍(long)(long)。暴(bao)龍(long)(long)亞(ya)(ya)科(ke)(ke)包(bao)含親緣關系(xi)較(jiao)(jiao)接近暴(bao)龍(long)(long),而離艾伯(bo)塔(ta)龍(long)(long)較(jiao)(jiao)遠(yuan)的(de)(de)物種;與艾伯(bo)塔(ta)龍(long)(long)亞(ya)(ya)科(ke)(ke)相比(bi),暴(bao)龍(long)(long)亞(ya)(ya)科(ke)(ke)的(de)(de)體格較(jiao)(jiao)重(zhong)型,頭顱骨的(de)(de)比(bi)例較(jiao)(jiao)大,以及較(jiao)(jiao)長的(de)(de)股骨。
勇士特暴龍最初被視為暴龍的一個種,某些近年的分類也支持這個說法。其他(ta)的(de)(de)(de)科(ke)學家則將它們列(lie)為獨立的(de)(de)(de)屬,并為暴(bao)龍(long)(long)(long)的(de)(de)(de)姐(jie)妹(mei)分類單元。在2003年(nian),一個親(qin)緣(yuan)分支(zhi)分類法研究(jiu)提出分支(zhi)龍(long)(long)(long)是(shi)特(te)暴(bao)龍(long)(long)(long)的(de)(de)(de)近親(qin),因(yin)為它們具(ju)有(you)(you)其他(ta)暴(bao)龍(long)(long)(long)亞(ya)(ya)(ya)科(ke)沒(mei)有(you)(you)的(de)(de)(de)頭部特(te)征(zheng)。如果(guo)屬實,將排除特(te)暴(bao)龍(long)(long)(long)是(shi)暴(bao)龍(long)(long)(long)的(de)(de)(de)一個異名(ming)的(de)(de)(de)可能性,并顯示暴(bao)龍(long)(long)(long)亞(ya)(ya)(ya)科(ke)在北美洲(zhou)與亞(ya)(ya)(ya)洲(zhou)演化(hua)出個別的(de)(de)(de)支(zhi)系。分支(zhi)龍(long)(long)(long)的(de)(de)(de)唯一標本具(ju)有(you)(you)幼年(nian)體(ti)的(de)(de)(de)特(te)征(zheng),但牙齒(chi)數量較多(duo),約76到78顆(ke),而且口鼻(bi)部上面(mian)有(you)(you)獨特(te)的(de)(de)(de)低矮骨質瘤(liu),因(yin)此(ci)并非(fei)特(te)暴(bao)龍(long)(long)(long)的(de)(de)(de)幼年(nian)體(ti)。
特暴龍(long)(long)(long)過去生存于潮濕的(de)泛(fan)濫(lan)平(ping)原,布滿者河道。在中國的(de)分布包括(kuo)黑龍(long)(long)(long)江,河南,山東(dong),廣東(dong),云(yun)南,內(nei)蒙(meng)古(gu)等地,國外主要(yao)是蒙(meng)古(gu)。在中國黑龍(long)(long)(long)江,河南,山東(dong),廣東(dong)的(de)所謂中國"暴龍(long)(long)(long)"其實都是特暴龍(long)(long)(long)。
特暴(bao)(bao)龍(long)(long)(long)是最(zui)(zui)大(da)型的暴(bao)(bao)龍(long)(long)(long)科(ke)動(dong)物之(zhi)一(yi)(yi)(yi),但略小于暴(bao)(bao)龍(long)(long)(long)。已知最(zui)(zui)大(da)型的個體(ti)(ti)身(shen)長(chang)12米(mi)(mi),頭部離地面約4.2米(mi)(mi)。一(yi)(yi)(yi)般體(ti)(ti)重(zhong)3噸(dun)到5噸(dun),最(zui)(zui)大(da)的化石(shi)體(ti)(ti)重(zhong)可達(da)到7.5噸(dun)。如同大(da)部分(fen)已知的暴(bao)(bao)龍(long)(long)(long)科(ke)恐(kong)龍(long)(long)(long),特暴(bao)(bao)龍(long)(long)(long)是種大(da)型、二足(zu)掠食動(dong)物,重(zhong)達(da)數(shu)噸(dun),擁(yong)有(you)數(shu)十(shi)顆大(da)型、銳利(li)的牙齒。特暴(bao)(bao)龍(long)(long)(long)的下頜(he)有(you)特殊(shu)的接合構造(zao)。另外,就(jiu)前(qian)肢/身(shen)體(ti)(ti)比例而言,特暴(bao)(bao)龍(long)(long)(long)擁(yong)有(you)暴(bao)(bao)龍(long)(long)(long)科(ke)中(zhong)最(zui)(zui)小型的前(qian)肢。特暴(bao)(bao)龍(long)(long)(long)是種大(da)型、二足(zu)掠食動(dong)物,重(zhong)達(da)數(shu)噸(dun)。特暴(bao)(bao)龍(long)(long)(long)唯一(yi)(yi)(yi)一(yi)(yi)(yi)個有(you)效種的體(ti)(ti)型:勇士特暴(bao)(bao)龍(long)(long)(long)Tarbosaurus bataar 8-12米(mi)(mi)。
暴龍科(ke)的身體(ti)外形差異不大(da)。特暴龍的頸部(bu)為S狀彎曲,其(qi)余的脊柱,包含尾巴,與地面(mian)保(bao)持者水平的姿態。
就前肢(zhi)/身體比例(li)而言,特暴龍擁有(you)暴龍科中最小型(xing)的前肢(zhi)。有(you)兩根迷你的手指(zhi)。后(hou)肢(zhi)長(chang)而粗厚(hou),將身體支撐(cheng)為二足的步態,上有(you)三(san)根腳(jiao)趾。長(chang)而重的尾巴可以平(ping)衡(heng)頭部(bu)與胸部(bu)的重量,將重心保持在臀部(bu)。
在(zai)2003年(nian),特暴(bao)(bao)龍(long)(long)(long)的(de)(de)顱(lu)(lu)骨(gu)(gu)(gu)(gu)首(shou)次(ci)經(jing)過完(wan)整的(de)(de)研究(jiu)。科學(xue)家們發現特暴(bao)(bao)龍(long)(long)(long)與(yu)北美洲暴(bao)(bao)龍(long)(long)(long)科之間有幾個顯著的(de)(de)差(cha)異(yi)。這(zhe)些差(cha)異(yi)與(yu)顱(lu)(lu)骨(gu)(gu)(gu)(gu)在(zai)咬合(he)時(shi),力量的(de)(de)傳遞(di)有關。當特暴(bao)(bao)龍(long)(long)(long)的(de)(de)上頜(he)(he)咬住物體時(shi),力量從上頜(he)(he)骨(gu)(gu)(gu)(gu)傳遞(di)到上頜(he)(he)周遭的(de)(de)顱(lu)(lu)骨(gu)(gu)(gu)(gu)。而北美洲暴(bao)(bao)龍(long)(long)(long)科咬合(he)時(shi),力量從上頜(he)(he)骨(gu)(gu)(gu)(gu)傳遞(di)到口鼻部(bu)上方的(de)(de)固定鼻骨(gu)(gu)(gu)(gu),然后再經(jing)由鼻骨(gu)(gu)(gu)(gu)與(yu)淚骨(gu)(gu)(gu)(gu)間的(de)(de)骨(gu)(gu)(gu)(gu)質連結,傳遞(di)到淚骨(gu)(gu)(gu)(gu)。
特暴龍的鼻骨(gu)(gu)(gu)與(yu)淚骨(gu)(gu)(gu)間(jian)缺乏骨(gu)(gu)(gu)質(zhi)的連結(jie)。但上(shang)頜(he)骨(gu)(gu)(gu)的后(hou)方有個大型突(tu)起,楔合入淚骨(gu)(gu)(gu)內;而北美洲暴龍的上(shang)頜(he)骨(gu)(gu)(gu)后(hou)突(tu)很小。這(zhe)個特征(zheng)顯(xian)示,咬合的力(li)量從特暴龍的上(shang)頜(he)骨(gu)(gu)(gu)直接(jie)傳遞到淚骨(gu)(gu)(gu)。而淚骨(gu)(gu)(gu)與(yu)額骨(gu)(gu)(gu)、前額骨(gu)(gu)(gu)之間(jian)更為牢固(gu)(gu)。由于上(shang)頜(he)骨(gu)(gu)(gu)、淚骨(gu)(gu)(gu)、額骨(gu)(gu)(gu)、前額骨(gu)(gu)(gu)之間(jian)牢牢地固(gu)(gu)定者,使得上(shang)頜(he)非(fei)常堅固(gu)(gu)。
另一個主要的差別則是堅固的下頜(he)。許多(duo)獸腳(jiao)類恐(kong)龍(long)(long),包(bao)括北美洲暴龍(long)(long)科在內,下頜(he)的齒(chi)骨與(yu)后面骨頭間有靈活(huo)的關節(jie)。特暴龍(long)(long)的隅骨側邊棱脊連(lian)接(jie)者齒(chi)骨后方的方形(xing)突,使它們(men)的下頜(he)無法靈活(huo)外內扳動。
有(you)些科學家提出假設,認為特(te)(te)(te)暴(bao)(bao)龍(long)(long)的(de)(de)堅硬頭(tou)部是種(zhong)適應演(yan)化,用來(lai)獵殺耐梅蓋特(te)(te)(te)組的(de)(de)大(da)型蜥腳類(lei)恐(kong)龍(long)(long),泰坦巨龍(long)(long)類(lei),因為晚(wan)白堊紀(ji)的(de)(de)北(bei)美洲并沒有(you)如此巨大(da)的(de)(de)恐(kong)龍(long)(long)存在(zai)。這種(zhong)頭(tou)部力學的(de)(de)差異也(ye)影(ying)響了暴(bao)(bao)龍(long)(long)科的(de)(de)種(zhong)系發生學。同樣發現(xian)于(yu)蒙古的(de)(de)分(fen)支龍(long)(long),也(ye)具(ju)有(you)類(lei)似(si)特(te)(te)(te)暴(bao)(bao)龍(long)(long)的(de)(de)頭(tou)部力學特(te)(te)(te)征,這顯示暴(bao)(bao)龍(long)(long)并非特(te)(te)(te)暴(bao)(bao)龍(long)(long)的(de)(de)近親(qin)。特(te)(te)(te)暴(bao)(bao)龍(long)(long)與(yu)暴(bao)(bao)龍(long)(long)的(de)(de)相似(si)處可(ke)能導因于(yu)它們巨大(da)的(de)(de)體型,是平行演(yan)化的(de)(de)結果。
一個在1948年發現(xian)的(de)顱(lu)骨(gu),一度被歸類于蛇(she)發女(nv)怪龍(long)的(de)G. lancinator,因(yin)為具有(you)顱(lu)腔(qiang),有(you)助于科學(xue)家了(le)(le)解(jie)特暴龍(long)的(de)腦部結(jie)構。在1965年,馬(ma)列夫(fu)制(zhi)作了(le)(le)一個石膏顱(lu)腔(qiang)模型,并(bing)做了(le)(le)初步的(de)腦部形狀檢(jian)查。在2005年,謝(xie)爾蓋·薩伐黎耶夫(fu)(Sergei V. Saveliev)制(zhi)作了(le)(le)一個聚氨酯(zhi)顱(lu)腔(qiang)模型,并(bing)做了(le)(le)更詳細的(de)腦部結(jie)構與功能(neng)研究。
根據暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)(de)(de)(de)(de)腦(nao)部(bu)結構研究,它們兩者的(de)(de)(de)(de)(de)腦(nao)部(bu)結構相似(si)(si),只有某些(xie)腦(nao)神(shen)經根部(bu)的(de)(de)(de)(de)(de)位置(zhi)不一樣,包含(han)三(san)叉神(shen)經與(yu)副神(shen)經。暴(bao)(bao)(bao)(bao)龍(long)(long)科(ke)的(de)(de)(de)(de)(de)腦(nao)部(bu)較類似(si)(si)鱷魚與(yu)其(qi)他爬(pa)蟲類,而較不類似(si)(si)鳥類。特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)(de)(de)(de)(de)腦(nao)部(bu)體(ti)積為184立方厘米。腦(nao)部(bu)的(de)(de)(de)(de)(de)大(da)型嗅(xiu)(xiu)球、末端(duan)神(shen)經、嗅(xiu)(xiu)神(shen)經,顯(xian)(xian)示特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)具(ju)有靈(ling)敏的(de)(de)(de)(de)(de)嗅(xiu)(xiu)覺(jue)(jue)(jue),這(zhe)點如同(tong)暴(bao)(bao)(bao)(bao)龍(long)(long)。犁鼻(bi)球大(da)型,而且(qie)與(yu)嗅(xiu)(xiu)球分離(li),顯(xian)(xian)示可以感應(ying)費(fei)洛蒙的(de)(de)(de)(de)(de)犁鼻(bi)器發展(zhan)(zhan)良(liang)好(hao)。這(zhe)個特(te)(te)征顯(xian)(xian)示特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)可能(neng)具(ju)有復雜的(de)(de)(de)(de)(de)求偶行為。聽(ting)(ting)(ting)神(shen)經也(ye)很(hen)大(da),顯(xian)(xian)示它們的(de)(de)(de)(de)(de)聽(ting)(ting)(ting)力很(hen)好(hao),可能(neng)用在聲音的(de)(de)(de)(de)(de)溝(gou)通與(yu)警告上。聽(ting)(ting)(ting)神(shen)經連(lian)接者發展(zhan)(zhan)良(liang)好(hao)的(de)(de)(de)(de)(de)前庭系(xi)統(tong),顯(xian)(xian)示它們的(de)(de)(de)(de)(de)平(ping)衡(heng)感與(yu)協調性很(hen)好(hao)。相反(fan)地,與(yu)視(shi)力有關的(de)(de)(de)(de)(de)腦(nao)部(bu)區域與(yu)神(shen)經較小。爬(pa)蟲類的(de)(de)(de)(de)(de)中腦(nao)頂蓋連(lian)接控制眼球的(de)(de)(de)(de)(de)視(shi)神(shen)經與(yu)動眼神(shen)經,是用來處理視(shi)力訊(xun)息,但特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)(de)(de)(de)(de)中腦(nao)頂蓋非常小。暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)(de)(de)(de)(de)眼睛朝前,因此(ci)具(ju)有一定程度的(de)(de)(de)(de)(de)立體(ti)視(shi)覺(jue)(jue)(jue);但特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)的(de)(de)(de)(de)(de)顱骨(gu)狹窄,眼睛朝向兩側,如同(tong)典型的(de)(de)(de)(de)(de)暴(bao)(bao)(bao)(bao)龍(long)(long)科(ke)動物(wu)。這(zhe)些(xie)特(te)(te)征顯(xian)(xian)示特(te)(te)暴(bao)(bao)(bao)(bao)龍(long)(long)較依靠嗅(xiu)(xiu)覺(jue)(jue)(jue)與(yu)聽(ting)(ting)(ting)覺(jue)(jue)(jue),而非視(shi)覺(jue)(jue)(jue)。
已知最大(da)(da)(da)型的特暴(bao)龍(long)(long)(long)顱(lu)骨長度(du)(du)超過(guo)1.3米(mi),小于暴(bao)龍(long)(long)(long)和諸(zhu)城暴(bao)龍(long)(long)(long)以及懼龍(long)(long)(long),但大(da)(da)(da)于其它的暴(bao)龍(long)(long)(long)科。如同暴(bao)龍(long)(long)(long),特暴(bao)龍(long)(long)(long)的顱(lu)骨高大(da)(da)(da),前段狹窄。顱(lu)骨后段擴張幅度(du)(du)不大(da)(da)(da),意味著特暴(bao)龍(long)(long)(long)所(suo)以不如暴(bao)龍(long)(long)(long)擁有(you)的立體視覺好(hao)。
顱(lu)骨的大型洞(dong)孔可減輕(qing)重量。特(te)暴龍的下頜(he)有(you)特(te)殊的接(jie)合(he)構(gou)造。
顎(e)部(bu)(bu)有60到64顆牙齒(chi)(chi),略少(shao)于暴(bao)龍(long),但大(da)于其(qi)它體型(xing)較(jiao)小的(de)暴(bao)龍(long)科(ke),例如蛇發(fa)(fa)女(nv)怪(guai)龍(long)與分支(zhi)龍(long)。大(da)部(bu)(bu)分的(de)牙齒(chi)(chi)橫剖面(mian)為橢圓形(xing),而(er)前上顎(e)骨(gu)的(de)牙齒(chi)(chi)橫剖面(mian)為D形(xing)。暴(bao)龍(long)科(ke)都具有這種異型(xing)齒(chi)(chi)特(te)征。上顎(e)骨(gu)的(de)牙齒(chi)(chi)最長,齒(chi)(chi)冠長達85公厘。如同其(qi)它大(da)型(xing)暴(bao)龍(long)科(ke)與現(xian)代的(de)科(ke)莫(mo)多龍(long),特(te)暴(bao)龍(long)的(de)幼年與近成(cheng)年個體的(de)身上發(fa)(fa)現(xian)過有齒(chi)(chi)痕,符合成(cheng)年個體或(huo)其(qi)它較(jiao)小型(xing)獸(shou)腳類(lei)恐龍(long)的(de)牙齒(chi)(chi)。
特暴龍(long)與分支龍(long)的下顎外側(ce)各有一道(dao)棱脊(ji)(ji),從(cong)隅(yu)骨延(yan)伸到齒骨后方,形成(cheng)相扣的結構。其它暴龍(long)科動物缺乏(fa)這道(dao)棱脊(ji)(ji),因此下顎更為靈活。
特暴龍位于食(shi)物鏈的頂端,是一種頂級掠食(shi)者,可能(neng)以大型恐(kong)龍為食(shi),例(li)如(ru)鴨嘴龍類的櫛(zhi)龍,或是蜥腳類的納摩蓋吐(tu)龍。
成年特暴(bao)(bao)龍(long)(long)(long)(long)(long)(long)(long)(long)可能(neng)與其它(ta)小型獸腳類恐龍(long)(long)(long)(long)(long)(long)(long)(long)有少許競(jing)爭(zheng)(zheng),例如傷(shang)齒龍(long)(long)(long)(long)(long)(long)(long)(long)科的(de)(de)(de)無聊龍(long)(long)(long)(long)(long)(long)(long)(long)、鴕鳥(niao)龍(long)(long)(long)(long)(long)(long)(long)(long)、蜥鳥(niao)龍(long)(long)(long)(long)(long)(long)(long)(long),以及偷蛋龍(long)(long)(long)(long)(long)(long)(long)(long)下目(mu)的(de)(de)(de)單足(zu)龍(long)(long)(long)(long)(long)(long)(long)(long)、瑞欽(qin)龍(long)(long)(long)(long)(long)(long)(long)(long),或者還有小掠龍(long)(long)(long)(long)(long)(long)(long)(long),一種有時被認為(wei)是基(ji)底暴(bao)(bao)龍(long)(long)(long)(long)(long)(long)(long)(long)超(chao)科的(de)(de)(de)恐龍(long)(long)(long)(long)(long)(long)(long)(long)。其它(ta)的(de)(de)(de)獸腳類恐龍(long)(long)(long)(long)(long)(long)(long)(long),包含:巨大的(de)(de)(de)鐮(lian)刀龍(long)(long)(long)(long)(long)(long)(long)(long)、似鳥(niao)龍(long)(long)(long)(long)(long)(long)(long)(long)下目(mu)的(de)(de)(de)似鵝龍(long)(long)(long)(long)(long)(long)(long)(long)、似雞龍(long)(long)(long)(long)(long)(long)(long)(long)、恐手(shou)龍(long)(long)(long)(long)(long)(long)(long)(long),鐮(lian)刀龍(long)(long)(long)(long)(long)(long)(long)(long)可能(neng)是草食性(xing),而上述似鳥(niao)龍(long)(long)(long)(long)(long)(long)(long)(long)類恐龍(long)(long)(long)(long)(long)(long)(long)(long)可能(neng)為(wei)雜(za)食性(xing),以小型動(dong)物為(wei)食,不會跟特暴(bao)(bao)龍(long)(long)(long)(long)(long)(long)(long)(long)競(jing)爭(zheng)(zheng)食物。
在(zai)2001年,布魯斯·羅斯柴爾德(Bruce Rothschild)等人發表一(yi)份(fen)獸腳類恐龍的(de)壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe)與肌(ji)腱(jian)撕裂(lie)傷研(yan)究,并研(yan)究它們(men)(men)的(de)行為模(mo)式。壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe)較(jiao)常(chang)導因(yin)(yin)于習慣性(xing)(xing)動作,較(jiao)少來自于外力(li)(li)沖擊。他們(men)(men)研(yan)究18個(ge)特(te)暴龍的(de)腳掌(zhang)骨(gu)(gu)骼,沒有(you)發現(xian)壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe)的(de)跡象;而(er)在(zai)10個(ge)被研(yan)究的(de)手(shou)掌(zhang)骨(gu)(gu)頭(tou)中(zhong),其(qi)中(zhong)一(yi)個(ge)手(shou)部(bu)骨(gu)(gu)頭(tou)有(you)發現(xian)壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe)的(de)跡象。腳部(bu)骨(gu)(gu)頭(tou)的(de)壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe),可以歸咎于奔(ben)跑、長(chang)途遷徙(xi)。手(shou)部(bu)骨(gu)(gu)頭(tou)的(de)壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe),極可能導因(yin)(yin)于捕(bu)抓獵物而(er)受傷。這些壓(ya)力(li)(li)性(xing)(xing)骨(gu)(gu)折(zhe)、肌(ji)腱(jian)撕裂(lie)傷跡象,顯示特(te)暴龍常(chang)因(yin)(yin)為捕(bu)抓獵物而(er)受傷,而(er)非吞(tun)食尸體。
在(zai)1946年(nian),一個蘇聯與蒙古挖(wa)掘團(tuan)隊在(zai)蒙古南戈壁省的(de)(de)耐梅(mei)蓋特組發現一個接(jie)近完整(zheng)的(de)(de)絕大部分大型頭顱骨(gu)與一些脊椎骨(gu),屬于一個大型的(de)(de)獸(shou)腳(jiao)類恐龍。
在(zai)1955年,蘇(su)聯(lian)古生物(wu)學家葉甫根尼·馬列(lie)夫(Evgeny Maleev)將這正模標本(ben)(PIN 551-1)建立為暴龍的一個種,勇士暴龍(Tyrannosaurus bataar)。種名(ming)баатар/baatar在(zai)蒙(meng)古語中意(yi)為"勇士",但被誤拼(pin)為bataar。
同(tong)年(nian),馬列夫(fu)將3個(ge)獸腳(jiao)類的化石敘(xu)述并命名,三者(zhe)都是頭(tou)顱骨(gu),外(wai)加部分身體,全是同(tong)一(yi)個(ge)挖(wa)掘團隊在1948到(dao)49年(nian)間發(fa)(fa)現(xian)的。第一(yi)個(ge)(編號PIN 551-2)被建立(li)為(wei)新(xin)屬(shu),埃夫(fu)雷莫夫(fu)特暴龍(long)(T. efremovi),ταρβο?/tarbos在希臘文(wen)文(wen)中意(yi)為(wei)"恐怖(bu)的"、"敬畏的",而σαυρο?/saurus意(yi)為(wei)"蜥蜴",種名是以(yi)蘇(su)聯古(gu)生物學家兼科幻小說作者(zhe)伊(yi)凡·埃夫(fu)雷莫夫(fu)(Ivan Yefremov)為(wei)名。另外(wai)兩個(ge)(編號PIN 553-1與(yu)PIN 552-2)被歸類于北美洲的蛇發(fa)(fa)女怪龍(long),G. lancinator與(yu)G. novojilovi。這三個(ge)標本都小于1946年(nian)發(fa)(fa)現(xian)的標本。
自從(cong)蘇聯與蒙古的挖(wa)掘(jue)(jue)團隊在(zai)(zai)40年代的挖(wa)掘(jue)(jue)過后,一個波(bo)蘭與蒙古的挖(wa)掘(jue)(jue)團隊再(zai)度(du)回到戈壁沙漠挖(wa)掘(jue)(jue),從(cong)1963年持續(xu)到1971年,發現(xian)了許(xu)多新的化石,并在(zai)(zai)耐梅蓋特(te)組發現(xian)了特(te)暴龍的新標(biao)本。
在(zai)1965年,蘇聯古生物(wu)學家阿納托利(li)·康斯(si)坦丁諾維奇·羅特杰(jie)斯(si)特文斯(si)基(Anatoly Konstantinovich Rozhdestvensky)認為馬列夫所發(fa)現的(de)標本(ben),其(qi)實是同(tong)種動物(wu)的(de)不同(tong)生長階段,而且不同(tong)于北美洲的(de)暴龍。
1979年,董枝明在(zai)(zai)根據欒(luan)川(chuan)縣發掘(jue)到五(wu)顆大(da)型牙齒命名(ming)為(wei)巒川(chuan)暴(bao)(bao)(bao)龍(long)(Tyrannosaurus luanchuanensis)和霸王龍(long)相似(si)種(zhong)(Tyrannosaurus cf. T. rex )的(de)動物后來(lai)就(jiu)被改為(wei)欒(luan)川(chuan)特(te)暴(bao)(bao)(bao)龍(long)等,而(er)欒(luan)川(chuan)特(te)暴(bao)(bao)(bao)龍(long)實(shi)際也(ye)(ye)不能成(cheng)立,因為(wei)后來(lai)的(de)研究把它(ta)歸入了勇士特(te)暴(bao)(bao)(bao)龍(long),認為(wei)它(ta)實(shi)際只(zhi)是(shi)勇士特(te)暴(bao)(bao)(bao)龍(long)的(de)青年個體。在(zai)(zai)欒(luan)川(chuan)特(te)暴(bao)(bao)(bao)龍(long)發現(xian)之前(qian),廣東河源發現(xian)過特(te)暴(bao)(bao)(bao)龍(long)未定種(zhong)的(de)化(hua)石(shi)。發現(xian)欒(luan)川(chuan)特(te)暴(bao)(bao)(bao)龍(long)的(de)秋(qiu)扒組巖(yan)石(shi),秋(qiu)扒組巖(yan)石(shi)是(shi)分布在(zai)(zai)潭頭(tou)盆地欒(luan)川(chuan)縣一(yi)帶的(de)白堊紀晚期地層,是(shi)由紫紅色的(de)砂泥巖(yan)構成(cheng),最底(di)部(bu)含有磚紅色的(de)礫石(shi),其他(ta)發現(xian)過特(te)暴(bao)(bao)(bao)龍(long)的(de)地層也(ye)(ye)都屬(shu)于(yu)白堊紀晚期。
在1986年,雖然(ran)已(yi)經發現(xian)許多(duo)特(te)暴(bao)龍(long)標本,但(dan)(dan)很少(shao)已(yi)確定的明確資料,特(te)暴(bao)龍(long)被推(tui)論(lun)與(yu)其他(ta)暴(bao)龍(long)科恐龍(long)有許多(duo)共同(tong)特(te)征。這些相近的特(te)征讓某些科學家(jia)提(ti)出當時(shi)的北(bei)美洲(zhou)與(yu)歐亞大陸之間有可能的連結(jie),也許是陸橋(qiao)。他(ta)將四個標本與(yu)新(xin)發現(xian)的化石,統(tong)合為勇士特(te)暴(bao)龍(long)(T.bataar)。后來的研(yan)究人員都同(tong)意(yi)羅特(te)杰斯特(te)文斯基的分類,包含馬列夫(fu)在內,但(dan)(dan)少(shao)數(shu)科學家(jia)使用(yong)埃夫(fu)雷莫夫(fu)特(te)暴(bao)龍(long),而非勇士特(te)暴(bao)龍(long)。
在(zai)1992年,美(mei)國古生物學家肯尼思·卡彭(peng)特(te)(Kenneth Carpenter)重新檢驗這(zhe)些化石,他根據頭顱(lu)骨(gu)的相似處,提出(chu)這(zhe)些化石大部分屬于暴(bao)龍(long),并將勇士(shi)特(te)暴(bao)龍(long)改回勇士(shi)暴(bao)龍(long);而蛇發女怪龍(long)的G. novojilovi是個較小型的暴(bao)龍(long)科(ke)動(dong)物,另外建立(li)為馬列夫龍(long)(Maleevosaurus novojilovi)。
在1995年(nian),喬治·奧利舍夫(fu)斯基(George Olshevsky)建立了(le)勇(yong)(yong)士成(cheng)(cheng)吉思汗龍(long)(long)(Jenghizkhan bataar),取(qu)代勇(yong)(yong)士特(te)暴龍(long)(long),并(bing)以成(cheng)(cheng)吉思汗為名;他并(bing)提出埃(ai)夫(fu)雷莫(mo)夫(fu)特(te)暴龍(long)(long)、馬(ma)列夫(fu)龍(long)(long)、勇(yong)(yong)士成(cheng)(cheng)吉思汗龍(long)(long)是三個獨(du)立的屬,都生(sheng)存于相同時期的耐梅蓋(gai)特(te)組(zu)。
在1999年(nian),一個研究(jiu)提出(chu)馬(ma)列(lie)夫龍(long)是(shi)特暴龍(long)的幼年(nian)個體。自從(cong)1999年(nian)以(yi)來,所有的研究(jiu)認為這(zhe)些(xie)化石皆為同種動物,勇士特暴龍(long)或是(shi)勇士暴龍(long)。
在1993年到1998年,日本與(yu)蒙古的挖(wa)掘團隊,以及21世紀(ji)初(chu),加拿大古生物學家菲力(li)·柯爾(Phil Currie)的挖(wa)掘小組,也發(fa)現了許多特暴龍化(hua)石。不算上私人收藏(zang)的標本,已發(fa)現了至少37個標本,包(bao)含超過15個完整和(he)部分的頭顱骨化(hua)石,與(yu)至少5個完整的顱后骨骸。
21世紀初(chu),廣東河源盆地也出土過大(da)型獸腳類的大(da)型暴(bao)龍(long)類牙齒(chi)化石,推測可(ke)能是特暴(bao)龍(long)。
特暴龍(long)的(de)(de)大部(bu)分(fen)化石(shi)(shi),是成(cheng)年(nian)或亞成(cheng)年(nian)個(ge)體(ti),很少發現(xian)幼(you)年(nian)個(ge)體(ti)的(de)(de)化石(shi)(shi)。在2006年(nian)發現(xian)的(de)(de)一個(ge)幼(you)年(nian)個(ge)體(ti)身體(ti)骨(gu)骼(ge),帶有完整的(de)(de)頭(tou)顱骨(gu),頭(tou)部(bu)長度為29厘米,使得古生物(wu)學家可以(yi)研究特暴龍(long)的(de)(de)生長模式(shi)。這個(ge)幼(you)年(nian)個(ge)體(ti)化石(shi)(shi),死亡時的(de)(de)年(nian)齡大約是2到3歲。與(yu)成(cheng)年(nian)個(ge)體(ti)相(xiang)比,這個(ge)幼(you)年(nian)頭(tou)顱骨(gu)的(de)(de)結構虛(xu)弱,牙(ya)齒較(jiao)細,顯示特暴龍(long)的(de)(de)幼(you)年(nian)個(ge)體(ti)、成(cheng)年(nian)個(ge)體(ti)占(zhan)據不同的(de)(de)生態(tai)位,以(yi)免競爭相(xiang)同食物(wu)來源(yuan)。