大(da)耳菊頭蝠(fu)(學名:Rhinolophus macrotis)是菊頭蝠(fu)科、菊頭蝠(fu)屬蝙(bian)蝠(fu)。體型較(jiao)小,耳特大(da),耳長(chang)23-27毫米。鼻葉(xie)(xie)之(zhi)馬蹄計寬大(da),中(zhong)間具(ju)(ju)明顯缺刻,前面兩(liang)側(ce)(ce)均(jun)具(ju)(ju)一小附葉(xie)(xie)。鞍狀葉(xie)(xie)寬約為其長(chang)之(zhi)半(ban),頂端呈(cheng)圓形,兩(liang)側(ce)(ce)緣(yuan)平行,基部(bu)擴大(da)并與鼻問葉(xie)(xie)聯成一線形杯狀葉(xie)(xie)。聯接葉(xie)(xie)特別低,呈(cheng)圓弧形。頂葉(xie)(xie)之(zhi)高(gao)度與鞍狀葉(xie)(xie)相似,側(ce)(ce)緣(yuan)微凹,頂端圓鈍。下唇具(ju)(ju)3條(tiao)縱(zong)行唇溝。前肢第(di)3掌(zhang)骨(gu)較(jiao)短,第(di)4、5掌(zhang)骨(gu)近乎等長(chang),第(di)3指(zhi)(zhi)(zhi)的第(di)2指(zhi)(zhi)(zhi)節長(chang)不及第(di)1指(zhi)(zhi)(zhi)節之(zhi)1.5倍,第(di)4、5指(zhi)(zhi)(zhi)的第(di)1指(zhi)(zhi)(zhi)節均(jun)略短于(yu)其第(di)2指(zhi)(zhi)(zhi)節。通體毛(mao)暗(an)灰(hui)色或(huo)灰(hui)褐色,基部(bu)灰(hui)白色,腹毛(mao)毛(mao)色淺(qian)淡。
大耳菊頭蝠(fu)(fu)習性生活(huo)于(yu)山洞中,數量不多,與其他(ta)蝙蝠(fu)(fu)共(gong)棲,一般停留在(zai)洞邊緣(yuan)或近洞口巖石頂壁上。以(yi)夜行性飛行生活(huo)的昆蟲為食。分布于(yu)孟(meng)加拉國、中國、印度、印度尼西(xi)亞(ya)(ya)(蘇門答臘(la))、老撾人民民主共(gong)和國、馬(ma)(ma)來西(xi)亞(ya)(ya)(馬(ma)(ma)來西(xi)亞(ya)(ya)半島)、緬甸、尼泊爾、巴基斯坦、泰國、越(yue)南、菲律賓。
存在不確定(ding):不丹(dan)
存在(zai):孟加(jia)拉國、中國、印(yin)度(du)(du)、印(yin)度(du)(du)尼(ni)西亞(ya)(蘇門答(da)臘)、老撾人民(min)民(min)主共和國、馬(ma)來(lai)西亞(ya)(馬(ma)來(lai)西亞(ya)半島)、緬甸(dian)、尼(ni)泊爾、巴基斯坦、泰國、越南(nan)、菲律賓。在(zai)中國分布(bu)于陜(shan)西、浙江、江西、貴州、四川(chuan)、福(fu)建、云(yun)南(nan)、廣西、湖(hu)南(nan)、山西。
體型較小(xiao),體重4.7(4-5)克,體長36.4(32-40.0)毫米,前(qian)臂長42.4(41-43.2)毫米。耳特(te)大,耳長23-27毫米。鼻葉(xie)(xie)之(zhi)馬蹄計(ji)寬大,中(zhong)間具明顯缺刻,前(qian)面(mian)兩(liang)側均具一小(xiao)附葉(xie)(xie)。鞍狀(zhuang)(zhuang)葉(xie)(xie)寬約為其長之(zhi)半,頂端呈圓(yuan)(yuan)(yuan)形(xing),兩(liang)側緣平行,基部擴(kuo)大并(bing)與(yu)鼻問(wen)葉(xie)(xie)聯(lian)成一線形(xing)杯狀(zhuang)(zhuang)葉(xie)(xie)。聯(lian)接葉(xie)(xie)特(te)別低(di),呈圓(yuan)(yuan)(yuan)弧形(xing)。頂葉(xie)(xie)之(zhi)高度與(yu)鞍狀(zhuang)(zhuang)葉(xie)(xie)相似,側緣微凹,頂端圓(yuan)(yuan)(yuan)鈍。下唇(chun)具3條(tiao)縱(zong)行唇(chun)溝(gou)。前(qian)肢第(di)(di)(di)3掌骨較短(duan),第(di)(di)(di)4、5掌骨近乎等(deng)長,第(di)(di)(di)3指的(de)第(di)(di)(di)2指節(jie)長不及第(di)(di)(di)1指節(jie)之(zhi)1.5倍,第(di)(di)(di)4、5指的(de)第(di)(di)(di)1指節(jie)均略(lve)短(duan)于(yu)其第(di)(di)(di)2指節(jie)。
通體(ti)毛(mao)暗灰(hui)色或灰(hui)褐色,基(ji)部灰(hui)白色,腹毛(mao)毛(mao)色淺淡。
頭(tou)骨(gu)吻部之(zhi)鼓泡(pao)發達(da)。矢狀脊不明顯,腭橋長度(du)超過上頰齒(chi)列長1/3。眶間距(ju)狹縮,顴寬(kuan)等于或(huo)略大予(yu)后頭(tou)寬(kuan)。下頜之(zhi)冠狀突(tu)低(di)。上頜第1前(qian)臼(jiu)齒(chi)較發達(da),位于齒(chi)列之(zhi)中。下頜第1前(qian)臼(jiu)齒(chi)位置偏向齒(chi)列外側。
大(da)耳菊(ju)頭蝠習性(xing)生活于(yu)山(shan)洞(dong)中,數量(liang)不多,與(yu)其他蝙蝠共棲,一(yi)般停留(liu)在洞(dong)邊緣或(huo)近洞(dong)口(kou)巖石頂壁上(shang)。以夜(ye)行性(xing)飛(fei)行生活的(de)昆(kun)蟲(chong)為食。
大(da)(da)耳(er)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus macrotis)屬philippinensis種(zhong)(zhong)(zhong)(zhong)組,Csorba等(deng)(2003)記錄了(le)該(gai)種(zhong)(zhong)(zhong)(zhong)組6種(zhong)(zhong)(zhong)(zhong)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu):大(da)(da)耳(er)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus macrotis)、馬(ma)氏菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus marshalli)、高鞍菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus paradoxolophus)、貴州菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus rex)、帝汶菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus montanus)和菲律賓菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus philippinensis)。Wu等(deng)(2008)發表(biao)philippinensis種(zhong)(zhong)(zhong)(zhong)組的一(yi)個新種(zhong)(zhong)(zhong)(zhong)——華南菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)(Rhinolophus huananus)。大(da)(da)耳(er)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)分(fen)布(bu)廣(guang),種(zhong)(zhong)(zhong)(zhong)內(nei)亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)分(fen)化較(jiao)多(duo),曾記載(zai)7個亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong),即:指明亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis macrotis)、印(yin)尼亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis dohrni)、菲律賓亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis hirsutus)、四(si)川亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis episcopus)、福建(jian)亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis caldwelli)、泰國(guo)亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis siamensis)和巴基斯坦亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis topali,Csorba&Bates,1995;Csorba et al.,2003;王應(ying)(ying)祥(xiang),2003)。大(da)(da)耳(er)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)泰國(guo)亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis siamensis)體形(xing)最小(xiao)(前臂(bei)長36.6毫米(mi),顱(lu)全(quan)長15.2毫米(mi)),Simmons(2005)將其(qi)獨(du)立成為一(yi)個種(zhong)(zhong)(zhong)(zhong)(Rhinolophus siamensis),并得到許(xu)多(duo)學者的支持(Wu et al.,2008;Smith,解(jie)焱(yan),2009)。因此,philippinensis種(zhong)(zhong)(zhong)(zhong)組應(ying)(ying)該(gai)包(bao)括8種(zhong)(zhong)(zhong)(zhong)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu),其(qi)中大(da)(da)耳(er)菊(ju)(ju)(ju)頭(tou)(tou)蝠(fu)種(zhong)(zhong)(zhong)(zhong)下可分(fen)為6個亞(ya)(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)。
大(da)耳(er)菊(ju)頭(tou)蝠(fu)在中國(guo)已(yi)記錄2個亞(ya)(ya)種(zhong):四川亞(ya)(ya)種(zhong)(Rhinolophus macrotis episcopus),分(fen)布(bu)在四川、重慶、貴州和陜西(xi)(xi);福(fu)建亞(ya)(ya)種(zhong)(Rhinolophus macrotis caldwelli),分(fen)布(bu)于浙江(jiang)、江(jiang)西(xi)(xi)、廣東、廣西(xi)(xi)、福(fu)建(Allen,1938-1940;羅(luo)蓉,1993;Smith,解焱,2009)。湖南發現的(de)大(da)耳(er)菊(ju)頭(tou)蝠(fu)的(de)前(qian)臂長、顱(lu)全長、顴(quan)寬、乳突外(wai)寬、上齒(chi)列長和下齒(chi)列長等(deng)測量數據與重慶萬縣及貴州開陽的(de)近(jin)似,而與福(fu)建亞(ya)(ya)種(zhong)(前(qian)臂長43.0毫米(mi),顱(lu)全長18.0毫米(mi))區別(bie)明顯(xian)(Allen,1938-1940;羅(luo)蓉,1993)。雖然(ran)華南菊(ju)頭(tou)蝠(fu)(Rhinolophus huananus)的(de)外(wai)形與大(da)耳(er)菊(ju)頭(tou)蝠(fu)非(fei)常相似,但前(qian)者(zhe)的(de)體形大(da)小(前(qian)臂長41.4毫米(mi),顱(lu)全長16.6毫米(mi))介于大(da)耳(er)菊(ju)頭(tou)蝠(fu)(Rhinolophus macrotis)和泰(tai)國(guo)大(da)耳(er)菊(ju)頭(tou)蝠(fu)(Rhinolophus siamensis)之間(Wu et al.,2008),易與之區別(bie)。因此將湖南湘西(xi)(xi)采(cai)集的(de)大(da)耳(er)菊(ju)頭(tou)蝠(fu)鑒(jian)定(ding)為四川亞(ya)(ya)種(zhong)(Rhinolophus macrotis episcopus)。
中文(wen)名(ming) 學(xue)名(ming) 命(ming)名(ming)者及年(nian)代
大耳(er)菊頭蝠福(fu)建亞種(zhong) Rhinolophus macrotis caldwelli Allen,1923
大(da)耳(er)菊頭蝠印尼亞種(zhong) Rhinolophus macrotis dohrni K. Andersen,1907
大耳菊(ju)頭蝠四川亞(ya)種 Rhinolophus macrotis episcopus Allen,1923
大耳(er)菊頭蝠菲律(lv)賓亞種 Rhinolophus macrotis hirsutus K. Andersen,1905
大耳菊頭蝠指(zhi)名亞種(zhong) Rhinolophus macrotis macrotis Blyth,1844
大耳菊頭蝠巴基斯坦亞種 Rhinolophus macrotis topali Csorba and Bates,1995
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