大(da)耳(er)菊頭(tou)蝠(學(xue)名(ming):Rhinolophus macrotis)是菊頭(tou)蝠科(ke)、菊頭(tou)蝠屬(shu)蝙(bian)蝠。體(ti)型較小,耳(er)特(te)大(da),耳(er)長(chang)(chang)23-27毫米。鼻(bi)葉(xie)之馬蹄計寬大(da),中間具(ju)(ju)(ju)明顯缺刻,前(qian)面兩側(ce)均具(ju)(ju)(ju)一小附葉(xie)。鞍(an)狀(zhuang)葉(xie)寬約為其(qi)長(chang)(chang)之半,頂(ding)端呈圓形(xing),兩側(ce)緣平行,基部(bu)擴大(da)并與(yu)鼻(bi)問葉(xie)聯(lian)成一線形(xing)杯狀(zhuang)葉(xie)。聯(lian)接葉(xie)特(te)別低,呈圓弧形(xing)。頂(ding)葉(xie)之高(gao)度與(yu)鞍(an)狀(zhuang)葉(xie)相(xiang)似(si),側(ce)緣微凹,頂(ding)端圓鈍。下唇具(ju)(ju)(ju)3條縱(zong)行唇溝。前(qian)肢第(di)(di)3掌骨較短,第(di)(di)4、5掌骨近乎等長(chang)(chang),第(di)(di)3指(zhi)(zhi)的第(di)(di)2指(zhi)(zhi)節(jie)長(chang)(chang)不及第(di)(di)1指(zhi)(zhi)節(jie)之1.5倍,第(di)(di)4、5指(zhi)(zhi)的第(di)(di)1指(zhi)(zhi)節(jie)均略(lve)短于其(qi)第(di)(di)2指(zhi)(zhi)節(jie)。通體(ti)毛暗(an)灰(hui)(hui)色(se)或(huo)灰(hui)(hui)褐色(se),基部(bu)灰(hui)(hui)白色(se),腹毛毛色(se)淺淡。
大耳菊頭(tou)蝠(fu)習性生(sheng)活(huo)于(yu)山洞中(zhong),數(shu)量不多(duo),與其他蝙蝠(fu)共棲(qi),一般停留在洞邊緣或近洞口巖石頂壁上。以夜行性飛(fei)行生(sheng)活(huo)的昆蟲為(wei)食。分布于(yu)孟(meng)加拉國、中(zhong)國、印度、印度尼(ni)西亞(蘇門(men)答(da)臘)、老(lao)撾人民民主共和國、馬(ma)來西亞(馬(ma)來西亞半島)、緬(mian)甸(dian)、尼(ni)泊爾(er)、巴基斯坦、泰(tai)國、越南(nan)、菲律賓。
存在不確定:不丹
存在(zai):孟(meng)加拉國(guo)、中國(guo)、印(yin)度、印(yin)度尼西亞(蘇(su)門答臘)、老(lao)撾人民(min)(min)民(min)(min)主共和國(guo)、馬來(lai)西亞(馬來(lai)西亞半島)、緬甸、尼泊爾、巴基斯坦、泰國(guo)、越南(nan)、菲律賓。在(zai)中國(guo)分(fen)布于陜西、浙江(jiang)、江(jiang)西、貴州、四川(chuan)、福(fu)建、云南(nan)、廣(guang)西、湖南(nan)、山西。
體(ti)型較(jiao)小,體(ti)重4.7(4-5)克,體(ti)長(chang)(chang)36.4(32-40.0)毫米,前臂長(chang)(chang)42.4(41-43.2)毫米。耳特大(da),耳長(chang)(chang)23-27毫米。鼻(bi)葉之(zhi)馬蹄計寬大(da),中(zhong)間具明顯(xian)缺刻,前面(mian)兩側均(jun)具一小附(fu)葉。鞍(an)狀(zhuang)葉寬約為其長(chang)(chang)之(zhi)半(ban),頂端(duan)呈圓(yuan)形(xing),兩側緣平(ping)行,基部擴大(da)并與(yu)(yu)鼻(bi)問葉聯成一線(xian)形(xing)杯(bei)狀(zhuang)葉。聯接(jie)葉特別低,呈圓(yuan)弧形(xing)。頂葉之(zhi)高度與(yu)(yu)鞍(an)狀(zhuang)葉相似,側緣微凹,頂端(duan)圓(yuan)鈍。下唇(chun)具3條縱(zong)行唇(chun)溝。前肢第(di)3掌骨較(jiao)短,第(di)4、5掌骨近(jin)乎等長(chang)(chang),第(di)3指(zhi)(zhi)(zhi)的(de)(de)第(di)2指(zhi)(zhi)(zhi)節(jie)長(chang)(chang)不及(ji)第(di)1指(zhi)(zhi)(zhi)節(jie)之(zhi)1.5倍,第(di)4、5指(zhi)(zhi)(zhi)的(de)(de)第(di)1指(zhi)(zhi)(zhi)節(jie)均(jun)略短于其第(di)2指(zhi)(zhi)(zhi)節(jie)。
通體毛(mao)暗灰(hui)色(se)或灰(hui)褐色(se),基部灰(hui)白(bai)色(se),腹毛(mao)毛(mao)色(se)淺淡(dan)。
頭骨吻部之(zhi)鼓泡發達。矢狀脊不明顯,腭橋長度超過上頰齒列長1/3。眶間距狹縮(suo),顴寬等(deng)于或(huo)略大予后頭寬。下(xia)頜之(zhi)冠狀突低。上頜第(di)(di)1前臼(jiu)齒較發達,位于齒列之(zhi)中。下(xia)頜第(di)(di)1前臼(jiu)齒位置(zhi)偏向齒列外(wai)側(ce)。
大耳菊頭蝠習性生(sheng)活(huo)于山洞(dong)中,數量不多,與(yu)其他(ta)蝙蝠共棲,一般(ban)停留(liu)在洞(dong)邊緣或近洞(dong)口巖(yan)石頂壁(bi)上。以(yi)夜行(xing)性飛行(xing)生(sheng)活(huo)的(de)昆蟲為食(shi)。
大(da)耳(er)菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus macrotis)屬philippinensis種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)組(zu),Csorba等(2003)記錄(lu)了該種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)組(zu)6種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)菊(ju)頭(tou)蝠(fu)(fu):大(da)耳(er)菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus macrotis)、馬氏菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus marshalli)、高鞍(an)菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus paradoxolophus)、貴州菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus rex)、帝(di)汶(wen)菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus montanus)和菲律(lv)賓菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus philippinensis)。Wu等(2008)發表philippinensis種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)組(zu)的一(yi)個(ge)新種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)——華(hua)南菊(ju)頭(tou)蝠(fu)(fu)(Rhinolophus huananus)。大(da)耳(er)菊(ju)頭(tou)蝠(fu)(fu)分(fen)(fen)布廣,種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)內(nei)亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)分(fen)(fen)化較多(duo)(duo),曾記載7個(ge)亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong),即:指明亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis macrotis)、印尼亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis dohrni)、菲律(lv)賓亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis hirsutus)、四川亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis episcopus)、福建亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis caldwelli)、泰(tai)國亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis siamensis)和巴基斯坦亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis topali,Csorba&Bates,1995;Csorba et al.,2003;王(wang)應(ying)祥,2003)。大(da)耳(er)菊(ju)頭(tou)蝠(fu)(fu)泰(tai)國亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus macrotis siamensis)體形最小(前(qian)臂長36.6毫(hao)米,顱(lu)全(quan)長15.2毫(hao)米),Simmons(2005)將其獨(du)立成為(wei)一(yi)個(ge)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)(Rhinolophus siamensis),并(bing)得到許多(duo)(duo)學者的支持(chi)(Wu et al.,2008;Smith,解焱,2009)。因此,philippinensis種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)組(zu)應(ying)該包括8種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)菊(ju)頭(tou)蝠(fu)(fu),其中大(da)耳(er)菊(ju)頭(tou)蝠(fu)(fu)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)下可分(fen)(fen)為(wei)6個(ge)亞(ya)(ya)(ya)(ya)種(zhong)(zhong)(zhong)(zhong)(zhong)(zhong)。
大(da)(da)耳(er)菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)在中(zhong)國已記(ji)錄2個(ge)亞(ya)(ya)(ya)種:四(si)川(chuan)亞(ya)(ya)(ya)種(Rhinolophus macrotis episcopus),分(fen)布(bu)在四(si)川(chuan)、重慶、貴州和陜(shan)西(xi);福建(jian)亞(ya)(ya)(ya)種(Rhinolophus macrotis caldwelli),分(fen)布(bu)于浙(zhe)江(jiang)、江(jiang)西(xi)、廣東、廣西(xi)、福建(jian)(Allen,1938-1940;羅蓉,1993;Smith,解焱(yan),2009)。湖(hu)南發(fa)現的(de)(de)大(da)(da)耳(er)菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)的(de)(de)前(qian)臂長(chang)(chang)(chang)、顱全長(chang)(chang)(chang)、顴寬、乳(ru)突外(wai)寬、上齒(chi)列長(chang)(chang)(chang)和下齒(chi)列長(chang)(chang)(chang)等測(ce)量數據(ju)與重慶萬縣(xian)及貴州開(kai)陽(yang)的(de)(de)近似,而與福建(jian)亞(ya)(ya)(ya)種(前(qian)臂長(chang)(chang)(chang)43.0毫(hao)米,顱全長(chang)(chang)(chang)18.0毫(hao)米)區(qu)別明顯(Allen,1938-1940;羅蓉,1993)。雖(sui)然華南菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)(Rhinolophus huananus)的(de)(de)外(wai)形(xing)與大(da)(da)耳(er)菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)非常(chang)相似,但(dan)前(qian)者(zhe)的(de)(de)體形(xing)大(da)(da)小(前(qian)臂長(chang)(chang)(chang)41.4毫(hao)米,顱全長(chang)(chang)(chang)16.6毫(hao)米)介于大(da)(da)耳(er)菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)(Rhinolophus macrotis)和泰(tai)國大(da)(da)耳(er)菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)(Rhinolophus siamensis)之(zhi)間(Wu et al.,2008),易與之(zhi)區(qu)別。因此(ci)將湖(hu)南湘西(xi)采集的(de)(de)大(da)(da)耳(er)菊(ju)頭(tou)(tou)(tou)(tou)蝠(fu)鑒定為四(si)川(chuan)亞(ya)(ya)(ya)種(Rhinolophus macrotis episcopus)。
中文名 學名 命名者及年代
大耳菊頭蝠福建亞種(zhong) Rhinolophus macrotis caldwelli Allen,1923
大耳(er)菊頭蝠印尼亞(ya)種 Rhinolophus macrotis dohrni K. Andersen,1907
大(da)耳(er)菊(ju)頭蝠四川亞種 Rhinolophus macrotis episcopus Allen,1923
大耳(er)菊頭蝠菲律(lv)賓亞種 Rhinolophus macrotis hirsutus K. Andersen,1905
大耳菊頭蝠(fu)指名亞種 Rhinolophus macrotis macrotis Blyth,1844
大耳菊頭蝠(fu)巴基斯坦(tan)亞種 Rhinolophus macrotis topali Csorba and Bates,1995
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