種(zhong)子成熟(shu)的(de)特點 如何辨別種(zhong)子是否(fou)成熟(shu)
種子成熟先是生理成熟,后是形態成熟。
生(sheng)理(li)(li)成(cheng)熟是指種(zhong)子營養物質(zhi)貯藏到一定程度,種(zhong)胚形成(cheng),種(zhong)實具(ju)有發(fa)芽能(neng)力(li)。生(sheng)理(li)(li)成(cheng)熟的(de)種(zhong)子含水(shui)量高,營養物質(zhi)處于易溶狀態,尚不(bu)能(neng)完全保(bao)護種(zhong)仁,不(bu)易防(fang)止水(shui)分散失。此時(shi)采(cai)集,種(zhong)仁急劇(ju)收縮(suo)不(bu)利于貯藏,會很快喪失發(fa)芽能(neng)力(li),抗逆性低,易受微生(sheng)物為害(hai)。但對深休(xiu)眠(mian)的(de)種(zhong)子,如山(shan)茱萸(yu)、山(shan)楂、椴樹、水(shui)曲柳(liu)等(deng),可用處于生(sheng)理(li)(li)成(cheng)熟期的(de)種(zhong)子,采(cai)后(hou)即播,以縮(suo)短(duan)休(xiu)眠(mian)期,提高發(fa)芽率。
形態(tai)成(cheng)(cheng)熟(shu)是指種實外部形態(tai)完(wan)全呈現出成(cheng)(cheng)熟(shu)特征(zheng),完(wan)成(cheng)(cheng)子胚發育過程,結束(shu)了(le)營養物(wu)質(zhi)的積累(lei),含水量降低,營養物(wu)質(zhi)轉化為難(nan)溶的脂肪、蛋白(bai)質(zhi)和淀粉,種子重量不再增加或增加很少(shao),呼吸作用微弱,種皮致密、堅實,抗逆性強(qiang),已進入休眠,耐(nai)貯藏。一(yi)般園林(lin)采(cai)(cai)種多是在形態(tai)成(cheng)(cheng)熟(shu)后采(cai)(cai)集(ji)。
一(yi)般樹種(zhong)(zhong)的(de)種(zhong)(zhong)子(zi)(zi)多是(shi)生(sheng)(sheng)理(li)(li)成(cheng)熟在先,一(yi)段時(shi)間之(zhi)后(hou)才達到形態(tai)成(cheng)熟。但也有的(de)種(zhong)(zhong)子(zi)(zi)生(sheng)(sheng)理(li)(li)與形態(tai)成(cheng)熟時(shi)間幾乎一(yi)致,或相隔時(shi)間很(hen)短(duan),如(ru)(ru)柳樹、榆樹、泡桐等,當種(zhong)(zhong)子(zi)(zi)達到生(sheng)(sheng)理(li)(li)成(cheng)熟后(hou)就自行脫落,應注意及時(shi)采收。也有生(sheng)(sheng)理(li)(li)后(hou)熟的(de)種(zhong)(zhong)子(zi)(zi),如(ru)(ru)銀杏(xing)。當假種(zhong)(zhong)皮(pi)呈黃(huang)色變軟,由樹上脫落,此時(shi)胚還未發育完(wan)全,不能立即播種(zhong)(zhong)。需(xu)經適當條件(jian)下的(de)貯(zhu)藏(zang),采取一(yi)定措施才能正常(chang)發芽。
同(tong)一地區不同(tong)的(de)樹種,由于生物學特性(xing)不同(tong),其種實(shi)成(cheng)熟(shu)(shu)時期(qi)也不同(tong)。大多(duo)數樹種的(de)種子成(cheng)熟(shu)(shu)在(zai)秋季,也有在(zai)春、夏(xia)季成(cheng)熟(shu)(shu),如柚子、鐵刀(dao)木、松(song)柏(bo)(bo)等(deng)早春成(cheng)熟(shu)(shu),楊樹、柳樹、榆(yu)樹等(deng)在(zai)春末夏(xia)初(chu)成(cheng)熟(shu)(shu),桑(sang)樹、柏(bo)(bo)樹等(deng)在(zai)夏(xia)季成(cheng)熟(shu)(shu);而苦楝、馬尾松(song)等(deng)入冬(dong)成(cheng)熟(shu)(shu)。
同(tong)(tong)一樹種(zhong)(zhong)在不同(tong)(tong)生(sheng)(sheng)長地(di)區、不同(tong)(tong)地(di)理位置,種(zhong)(zhong)實(shi)(shi)的(de)成(cheng)熟(shu)期也不同(tong)(tong)。一般生(sheng)(sheng)長在南方的(de)樹種(zhong)(zhong)比生(sheng)(sheng)長在北方的(de)種(zhong)(zhong)實(shi)(shi)成(cheng)熟(shu)早。
同一(yi)樹種(zhong)(zhong)雖生(sheng)(sheng)長在(zai)同一(yi)地區(qu),但由于(yu)立地條件、天氣變化等差異,種(zhong)(zhong)子(zi)成熟(shu)(shu)期也不(bu)同。生(sheng)(sheng)于(yu)沙質土(tu)壤比(bi)粘質土(tu)壤樹種(zhong)(zhong)種(zhong)(zhong)子(zi)的(de)成熟(shu)(shu)要早(zao),陽坡比(bi)陰坡的(de)成熟(shu)(shu)要早(zao),林緣比(bi)林內的(de)成熟(shu)(shu)要早(zao),高溫(wen)干旱(han)地區(qu)年份(fen)比(bi)冷涼多雨地區(qu)成熟(shu)(shu)要早(zao)。
種子成(cheng)(cheng)熟(shu)特(te)征可(ke)分為3類:漿果(guo)類成(cheng)(cheng)熟(shu)期(qi)果(guo)皮變軟,顏色由綠變紅、黃、紫色等,并具有香味,多能自(zi)行脫落,應注意及時采摘。
干果(guo)類(lei)(莢果(guo)、蒴果(guo)、翅果(guo)等)成熟時果(guo)皮變為褐色,干燥開裂,也(ye)有(you)在樹枝上(shang)宿存的,如(ru)刺槐、紫藤(teng)、烏(wu)桕、衛矛類(lei)等。
球果類果鱗(lin)干裂(lie)、硬化、變色,種鱗(lin)開裂(lie)散出種子,如油松(song)、白皮松(song)、馬(ma)尾松(song)、華山松(song)。